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      <FootnoteTextInd>X</FootnoteTextInd>
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      <SupplementalInformationDetail>
        <FormAndLineReferenceDesc>FORM 990, PART III, LINE 1</FormAndLineReferenceDesc>
        <ExplanationTxt>PUBLIC LIBRARY OF SCIENCE (PLOS) IS A NONPROFIT PUBLISHER AND ADVOCACY ORGANIZATION FOUNDED TO ACCELERATE PROGRESS IN SCIENCE AND MEDICINE BY LEADING A TRANSFORMATION IN RESEARCH COMMUNICATION. PLOS IS A LEADER IN OPEN ACCESS PUBLISHING WITH A SUITE OF SEVEN INFLUENTIAL JOURNALS, PROVIDING TENS OF THOUSANDS OF RESEARCH ARTICLES THAT ARE IMMEDIATELY FREE TO ACCESS ONLINE, DISTRIBUTE AND REUSE. SHARING RESEARCH ENCOURAGES PROGRESS, FROM PROTECTING THE BIODIVERSITY OF THE PLANET TO FINDING MORE EFFECTIVE TREATMENTS FOR DISEASES SUCH AS CANCER AND MALARIA. THE ORGANIZATION'S CORE OBJECTIVES ARE TO: *PROVIDE WAYS TO OVERCOME UNNECESSARY BARRIERS TO IMMEDIATE AVAILABILITY, ACCESS AND USE OF RESEARCH *PURSUE A PUBLISHING STRATEGY THAT OPTIMIZES THE QUALITY AND INTEGRITY OF THE PUBLICATION PROCESS *DEVELOP INNOVATIVE APPROACHES TO THE ASSESSMENT, ORGANIZATION AND REUSE OF IDEAS AND DATA PLOS SUPPORTS OPEN ACCESS TO MAKE SCIENTIFIC ARTICLES IMMEDIATELY AND FREELY AVAILABLE TO ANYONE, ANYWHERE FOR THEM TO DOWNLOAD, PRINT, DISTRIBUTE, READ, AND REUSE WITHOUT CHARGE OR OTHER RESTRICTIONS, AS LONG AS THE AUTHOR IS PROPERLY ATTRIBUTED. OPEN ACCESS FUELS DISCOVERY BY FREEING RESEARCH FROM ITS BARRIERS. A BROAD COMMUNITY BENEFITS FROM THE WORK OF PLOS AS AN OPEN ACCESS INNOVATOR: SCIENTISTS MORE RAPIDLY BUILD ON THE WORK OF OTHERS IN THEIR FIELD, LIBRARIANS ENABLE WIDER DISSEMINATION OF PUBLISHED WORK, FUNDERS MAXIMIZE THEIR RETURN ON INVESTMENT IN THE RESEARCH THEY FUND, ENTREPRENEURS ACCESS ARTICLES TO DEVELOP NEW TECHNOLOGIES, RESEARCH INSTITUTIONS ACT ON THEIR DIRECTIVES AND THE PUBLIC BENEFITS FROM SCIENTIFIC AND MEDICAL ADVANCES.</ExplanationTxt>
      </SupplementalInformationDetail>
      <SupplementalInformationDetail>
        <FormAndLineReferenceDesc>FORM 990, PART III, LINE 4A</FormAndLineReferenceDesc>
        <ExplanationTxt>AN INNOVATIVE AND INFLUENTIAL VENUE FOR RESEARCH AND COMMENT ON THE MAJOR CHALLENGES TO HUMAN HEALTH WORLDWIDE, PLOS MEDICINE PUBLISHES ARTICLES ACROSS ALL AREAS OF MEDICAL SCIENCE, CLINICAL PRACTICE AND HEALTH POLICY THAT HAVE THE POTENTIAL TO DIRECTLY AND SUBSTANTIALLY INFORM CLINICAL PRACTICE OR HEALTH POLICY, OR TO PROVIDE SUBSTANTIAL, NOVEL MECHANISTTHERE WERE SEVERAL NOTABLE MILESTONES THIS YEAR AS PLOS CELEBRATED ITS FIRST DECADE AS AN OPEN ACCESS PUBLISHER AND WRAPPED THE YEAR BY PUBLISHING ITS 100,000TH ARTICLE. KEY MILESTONES FOR EACH JOURNAL SHOWCASED THE QUALITY, DIVERSITY AND GLOBAL REACH IMPACT OF OFFERINGS BY PLOS JOURNALS. FOR EXAMPLE, PLOS BIOLOGY CELEBRATED THE TEN-YEAR ANNIVERSARY OF ITS FIRST ARTICLES, AND PLOS COMPUTATIONAL BIOLOGY'S 'TEN SIMPLE RULES' SERIES OF EDITORIALS ON PROFESSIONAL DEVELOPMENT REACHED ONE MILLION PAGE VIEWS. THE ORGANIZATION ALSO WELCOMED NEW LEADERSHIP WITH ELIZABETH MARINCOLA AND RICHARD HEWITT JOINING PLOS AS NEW CEO AND CFO, RESPECTIVELY. PLOS CONTINUES ITS EFFORTS TO DRIVE OPENNESS IN DATA USE AND RE-USE WITH THE GOAL FOR ANYONE TO BE ABLE TO ANALYZE DATA (TO REPEAT AUTHORS' RESULTS) AND REUSE DATA (TO BUILD ON THE AUTHORS' FINDINGS). IN COLLABORATION WITH 27 SPONSORING ORGANIZATIONS THAT SHARE A COMMON BELIEF IN THE TRANSFORMATIVE IMPACT OF OPEN ACCESS, INCLUDING MAJOR SPONSORS WELLCOME TRUST AND GOOGLE, PLOS LAUNCHED THE ACCELERATING SCIENCE AWARD PROGRAM IN MAY 2013. THE PROGRAM ATTRACTED OVER 200 NOMINATIONS FROM MORE THAN 30 COUNTRIES WITH THE TOP THREE WINNERS AWARDED A PRIZE OF $30,000 EACH DURING OCTOBER'S OPEN ACCESS WEEK. THE ASAP AWARD RECOGNIZES INDIVIDUALS WHO HAVE APPLIED SCIENTIFIC RESEARCH--PUBLISHED THROUGH OPEN ACCESS--TO INNOVATE IN ANY FIELD AND BENEFIT SOCIETY. THE ASAP WEBSITE IS THE ARCHIVAL LOCATION OF FINAL PROGRAM MATERIALS FEATURING MULTIMEDIA CONTENT THAT INCLUDES: THE ACCELERATING IMPACT AWARD PROGRAM VIDEO DESCRIBING THE AWARD PROGRAM AND CONTAINING COMMENTS FROM ADVOCATES IN PUBLISHING, POLICY, ACADEMIA AND INDUSTRY; THE INDIVIDUAL FINALIST VIDEOS DESCRIBING THE AWARD-WINNING INNOVATIVE WORK THAT HIGHLIGHTS REAL-WORLD APPLICATIONS AND ADVANTAGES OF OPEN ACCESS RESEARCH AND USE OF OPEN ACCESS RESOURCES; AND A PROGRAM PORTFOLIO BOOKLET, PUBLISHED IN CONJUNCTION WITH ASAP, THAT DETAILS THE HONOREES AND THE BENEFITS OF OPEN ACCESS, INCLUDING EXAMPLES OF HOW OPEN ACCESS PUBLISHING IS AN ECONOMIC ENGINE FOR JOBS AND NEW BUSINESSES. OVER THE PAST YEAR PLOS CONTINUED ITS PROGRAM WITH DRYAD DIGITAL REPOSITORY. DRYAD HELPS PLOS AUTHORS MAKE THEIR DATA RE-USEABLE AND ACCESSIBLE BY PROVIDING A PERMANENT OPEN ACCESS REPOSITORY AND ASSIGNING UNIQUE DIGITAL OBJECT IDENTIFIERS, DOIS, FOR EACH DATA PACKAGE, COMPRISING ALL THE DATA ASSOCIATED WITH A SINGLE PUBLISHED ARTICLE. TWO PLOS JOURNALS, PLOS BIOLOGY AND PLOS GENETICS, HAVE USED THE DRYAD REPOSITORY TO HOST DATA WHILE ARTICLES ARE UNDER REVIEW, PROVIDING CONFIDENTIAL ACCESS TO THE UNDERLYING DATA FOR EDITORS AND PEER-REVIEWERS. MEANWHILE, PLOS IS WORKING TO EVOLVE THE SCIENTIFIC ARTICLE ITSELF INTO A MORE DYNAMIC, LIVING FORM OF COMMUNICATION. PLOS ESTABLISHED A PARTNERSHIP WITH FIGSHARE TO MAKE THE DATA PUBLISHED IN OUR ARTICLES MORE ACCESSIBLE AND DISCOVERABLE. THE FIGSHARE WIDGET DISPLAYS THE CONTENTS OF SUPPORTING INFORMATION FILES WITHIN THE ARTICLE, REGARDLESS OF FILE TYPE (WHETHER DATASET, TEXT, DOCUMENTS, ANIMATED, VIDEO, OR PRESENTATION FILES). THIS ALLOWS USERS TO SEARCH, MAGNIFY AND DOWNLOAD THE MATERIAL IN WAYS NOT PREVIOUSLY SUPPORTED. THESE DEVELOPMENTS COINCIDE WITH THE COMPLETION OF PHASE TWO OF THE FULL RE-DESIGN OF THE PLOS WEBSITES. PHASE ONE FOR THE REDESIGN OF THE PLOS JOURNAL WEBSITES WAS COMPLETED LAST YEAR. THIS PROJECT HAD THREE PRIMARY GOALS: TO ENSURE READERS CAN QUICKLY ASSESS THE RELEVANCE AND IMPORTANCE OF AN ARTICLE; TO IMPROVE SITE NAVIGATION; AND TO LAUNCH A FLEXIBLE PLATFORM FROM WHICH TO BUILD OUT FUTURE INNOVATIONS. THE NEW WEBSITES OFFER MORE PROMINENT FIGURES, AN IMMEDIATELY ACCESSIBLE SUBSET OF ARTICLE-LEVEL METRICS (ALM) DATA CALLED METRICS SIGNPOSTS, CUSTOM SAVED SEARCH, AUTHOR DATA, FASTER NAVIGATION, AND MUCH MORE. ADDITIONALLY, ENHANCEMENTS TO ARTICLE PAGES ENABLES THE ADDITION OF MEDIA STORIES TO INDIVIDUAL ARTICLE PAGES. PLOS WAS AWARDED 'BEST IN CLASS' FOR NONPROFIT WEBSITES AT THE 2013 INTERACTIVE MEDIA AWARDS. PLOS ENCOURAGES THE EVALUATION OF RESEARCH AT THE ARTICLE LEVEL. PLOS WAS ONE OF THE FIRST PUBLISHERS TO PLACE ARTICLE-LEVEL METRICS ON EVERY ARTICLE. ALMS PROVIDE A FULL AND CONSTANTLY UPDATED PICTURE OF HOW A PAPER IS BEING USED FROM THE MOMENT IT IS PUBLISHED ONLINE AND INCLUDE TRADITIONAL METRICS (SUCH AS CITATIONS AND ONLINE USAGE) ALONGSIDE MORE IMMEDIATE ALTERNATIVE METRICS SUCH AS COVERAGE ON SOCIAL MEDIA. PLOS ALMS PROVIDE METRICS ACROSS A SUITE OF SOURCES THAT INCLUDES HTML, XML, AND PDF DOWNLOADS, PUBMED CENTRAL USE, SCOPUS, WEB OF SCIENCE AND CROSSREF CITATIONS, CITUULIKE AND MENDELEY REFERENCE MANAGEMENT SERVICES, RESEARCHBLOGGING.ORG AND NATURE BLOGS, AND SOCIAL MEDIA PLATFORMS TWITTER AND FACEBOOK. MANY OTHER PUBLISHERS HAVE SINCE ADOPTED THE IDEA AND NEW BUSINESSES ARE NOW EMERGING TO DEMONSTRATE RESEARCH IMPACT IN THESE WAYS. OVER THE LAST YEAR, PLOS UPDATED ITS ALM PROGRAM TO ALLOW USERS TO VIEW AND DOWNLOAD ALMS FOR ANY ARTICLE. THESE ALM REPORTS ENABLE USERS TO CHART THE DATA AND IDENTIFY PATTERNS AND TRENDS FOR FURTHER DISCUSSION. PLOS ALSO RELEASED THE FIRST ITERATION OF ALM RELATIVE METRICS TO PROVIDE CONTEXT: THEY DISPLAY THE AVERAGE USAGE CALCULATION FOR SIMILAR ARTICLES PUBLISHED IN THE SAME TIME PERIOD AND RESEARCH AREAS, ALLOWING USERS TO EVALUATE RELATIVE PERFORMANCE IN DIFFERENT FIELDS. PLOS HAS USED THE CREATIVE COMMONS ATTRIBUTION (CC BY) LICENSE FOR ALMOST 10 YEARS AS THE DEFAULT FOR THE RESEARCH THAT IT PUBLISHES. PLOS ADOPTED CC BY V4.0 BEGINNING IN MID-DECEMBER FOR PLOS ONE AND FROM JANUARY 1, 2014 FOR ALL OTHER PLOS JOURNALS. THE NEW LICENSES ARE THE RESULT OF AN OPEN COMMUNITY PROCESS WITH STAKEHOLDERS FROM A WIDE RANGE OF DOMAINS, INCLUDING RESEARCH, EDUCATION, AND THE CREATIVE ARTS, AND PLOS IS PROUD TO HAVE BEEN INVOLVED IN THE EFFORT TO MAKE THE LICENSES WORK FOR RESEARCHERS. AN IMPORTANT ASPECT OF THE NEW LICENSES IS THAT THEY COMBINE THE EXPERIENCE OF SEVERAL YEARS OF DEVELOPING LOCALIZED LICENSE VARIANTS INTO ONE INTERNATIONAL LICENSE, ENSURING GLOBAL COMPATIBILITY AND EASE OF USE FOR ALL RESEARCHERS, WHEREVER THEY MAY BE BASED. TWO ASPECTS OF THE VERSION 4.0 LICENSES ARE PARTICULARLY IMPORTANT FOR RESEARCHERS SINCE THEY ADDRESS ISSUES THAT COULD HAVE MADE REUSE OF PUBLISHED RESEARCH MORE CUMBERSOME. FIRSTLY, THE REUSE RIGHTS FOR DATA WITHIN AN ARTICLE ARE MADE CLEARER AND MORE CONSISTENT BETWEEN DIFFERENT COUNTRIES AND REGIONS. SECOND, THE LICENSES PROVIDE FLEXIBILITY ON ATTRIBUTION. THIS IS IMPORTANT FOR RESEARCH, AND PARTICULARLY FOR TEXT AND DATA MINING, WHERE A MULTITUDE OF ARTICLES MIGHT BE ANALYZED TOGETHER. THE NEW LICENSES STILL ABSOLUTELY REQUIRE ATTRIBUTION BUT ALLOW ALL ATTRIBUTIONS TO A LARGE CORPUS TO BE COLLECTED TOGETHER. PLOS CONTINUES TO COLLABORATE WITH OTHERS IN THE OPEN ACCESS COMMUNITY TO CAPTURE THE INTEREST AND SUPPORT OF RESEARCHERS, GOVERNMENTS AND FUNDERS. PLOS PARTNERS WITH OTHER LIKE-MINDED NONPROFITS AND UNIVERSITIES TO EDUCATE THE PUBLIC ABOUT POLICIES THAT IMPACT SCIENCE RESEARCH DISSEMINATION AS PART OF ITS ADVOCACY EFFORTS. THESE COMMUNITY ENGAGEMENTS HAVE YIELDED POSITIVE IMPACT IN SHAPING POLICY CONVERSATIONS IN THE US, UK AND EU FOR TAX FUNDED RESEARCH PAPERS TO BE MADE ACCESSIBLE TO THE PUBLIC. BEING FREELY ABLE TO READ CONTENT IS ONLY THE BEGINNING. PLOS IS ALSO FOCUSED ON THE BROADER RE-USE VALUE OF OPEN CONTENT AND HAS COLLABORATED WITH THE SCHOLARLY PUBLISHING AND ACADEMIC RESOURCES COALITION, SPARC, AND OPEN ACCESS SCHOLARLY PUBLISHERS ASSOCIATION, OASPA, TO ENABLE DEEPER UNDERSTANDING AND COMPARISON OF PUBLISHERS' OPEN ACCESS POLICIES. TOGETHER, THE ORGANIZATIONS HAVE CREATED "HOWOPENISIT?" A GUIDE THAT IDENTIFIES THE CORE COMPONENTS OF OPEN ACCESS, THEIR CHARACTERISTICS AND HOW THEY ARE IMPLEMENTED ACROSS THE SPECTRUM FROM OPEN ACCESS TO CLOSED ACCESS. THIS THEME OF "INFORMED DECISIONS" IS ALSO REFLECTED IN PLOS' ONGOING ENDORSEMENT OF ALMS. PLOS ALSO PROVIDED $3.4 MILLION IN THE FORM OF PARTIAL OR FULL WAIVERS TO AUTHORS AS PART OF OUR PUBLICATION FEE ASSISTANCE. AS A NONPROFIT PUBLISHER, ONE CORE OBJECTIVE IS TO PROVIDE WAYS TO OVERCOME UNNECESSARY BARRIERS TO IMMEDIATE AVAILABILITY, ACCESS AND USE OF RESEARCH, WHICH AT TIMES MAY INCLUDE COST. THOUGH THIS AMOUNT APPEARS AS A CONTRA-REVENUE FOR FINANCIAL REPORTING PURPOSES, THIS IS A MAJOR PROGRAM EXPENSE FOR PLOS WHICH DEMONSTRATES THE ORGANIZATION'S SUPPORT TO SCIENCE AND GLOBAL PUBLIC COMMUNITY. MUCH OF THE HARDSHIP SUPPORT GOES TO AUTHORS FROM EMERGING ECONOMIES AND EARLY CAREER SCIENTISTS IN DEVELOPED ECONOMIES WHO HAVE BEEN UNSUCCESSFUL IN SECURING ADEQUATE FUNDING SUPPORT.</ExplanationTxt>
      </SupplementalInformationDetail>
      <SupplementalInformationDetail>
        <FormAndLineReferenceDesc>FORM 990, PART VI, SECTION B, LINE 11</FormAndLineReferenceDesc>
        <ExplanationTxt>FORM 990 IS PREPARED BY AN OUTSIDE TAX PROFESSIONAL WITH ASSISTANCE FROM THE ORGANIZATION. THE FORM IS THEN REVIEWED BY THE CFO AND MEMBERS FROM THE AUDIT COMMITTEE. AFTER REVIEW AND MODIFICATIONS WHERE NECESSARY, THE FINAL VERSION OF THE TAX RETURN IS PROVIDED TO THE BOARD OF DIRECTORS. THE CFO SIGNS AND FILES ALL REQUIRED TAX FILINGS.</ExplanationTxt>
      </SupplementalInformationDetail>
      <SupplementalInformationDetail>
        <FormAndLineReferenceDesc>FORM 990, PART VI, SECTION B, LINE 12C</FormAndLineReferenceDesc>
        <ExplanationTxt>THE GENERAL COUNSEL AND SECRETARY OVERSEES THE ANNUAL CONFLICT OF INTEREST DISCLOSURE PROCESS. A COMMITTEE OF THE BOARD OF DIRECTORS REVIEWS ALL POTENTIAL CONFLICTS OF INTEREST AT LEAST ANNUALLY. THE CEO AND ALL BOARD MEMBERS ARE REQUIRED TO DISCLOSE (IN WRITING) POTENTIAL CONFLICTS AND ANY RELATED PARTY AFFILIATIONS. LOANS BETWEEN THE ORGANIZATION AND MEMBERS OF MANAGEMENT AND THE BOARD ARE STRICTLY PROHIBITED. THE ORGANIZATION SEEKS FULL TRANSPARENCY ON ALL RELATIONSHIPS. ANY POTENTIAL CONFLICTS (IN FACT OR APPEARANCE) ARE DISCUSSED OPENLY AND RESOLVED IN ACCORDANCE WITH THE ORGANIZATION'S POLICIES AND PROCEDURES.</ExplanationTxt>
      </SupplementalInformationDetail>
      <SupplementalInformationDetail>
        <FormAndLineReferenceDesc>FORM 990, PART VI, SECTION B, LINE 15</FormAndLineReferenceDesc>
        <ExplanationTxt>THE COMPENSATION COMMITTEE OF THE BOARD OF DIRECTORS REVIEWS THE COMPENSATION OF ALL HIGH-LEVEL PERSONNEL ANNUALLY IN ACCORDANCE WITH IRS RULES AND REGULATIONS. EFFORTS ARE MADE TO SECURE COMPENSATION DATA FROM INDUSTRY SOURCES IN ORDER TO DETERMINE COMPETITIVENESS AND APPROPRIATENESS OF SALARIES AND BENEFITS. EVERY EFFORT IS MADE TO ENSURE THAT THE PROCESS IS THOROUGH AND TRANSPARENT IN ACCORDANCE WITH APPROPRIATE GOVERNANCE AND REGULATORY COMPLIANCE. THIS IS PERFORMED ANNUALLY AND THE PROCESS IS MANAGED BY THE HR DIRECTOR. REVIEW ENCOMPASSES CEO, CFO, EXECUTIVES AND KEY EMPLOYEES.</ExplanationTxt>
      </SupplementalInformationDetail>
      <SupplementalInformationDetail>
        <FormAndLineReferenceDesc>FORM 990, PART VI, SECTION C, LINE 19</FormAndLineReferenceDesc>
        <ExplanationTxt>PLOS MAKES ITS GOVERNING DOCUMENTS, CONFLICT OF INTEREST POLICY, AND FINANCIAL STATEMENTS AVAILABLE TO THE PUBLIC. THESE DOCUMENTS ARE ALSO POSTED ON OUR WEBSITE.</ExplanationTxt>
      </SupplementalInformationDetail>
    </IRS990ScheduleO>
  </ReturnData>
</Return>