﻿<?xml version="1.0" encoding="utf-8"?>
<Return xmlns="http://www.irs.gov/efile" xmlns:xsi="http://www.w3.org/2001/XMLSchema-instance" xsi:schemaLocation="http://www.irs.gov/efile" returnVersion="2024v5.2">
  <ReturnHeader binaryAttachmentCnt="0">
    <ReturnTs>2025-11-17T10:01:49-06:00</ReturnTs>
    <TaxPeriodEndDt>2024-12-31</TaxPeriodEndDt>
    <PreparerFirmGrp>
      <PreparerFirmEIN>946214841</PreparerFirmEIN>
      <PreparerFirmName>
        <BusinessNameLine1Txt>ARMANINO ADVISORY LLC</BusinessNameLine1Txt>
      </PreparerFirmName>
      <PreparerUSAddress>
        <AddressLine1Txt>2700 CAMINO RAMON STE 350</AddressLine1Txt>
        <CityNm>SAN RAMON</CityNm>
        <StateAbbreviationCd>CA</StateAbbreviationCd>
        <ZIPCd>945835004</ZIPCd>
      </PreparerUSAddress>
    </PreparerFirmGrp>
    <ReturnTypeCd>990</ReturnTypeCd>
    <TaxPeriodBeginDt>2024-01-01</TaxPeriodBeginDt>
    <Filer>
      <EIN>680492065</EIN>
      <BusinessName>
        <BusinessNameLine1Txt>PUBLIC LIBRARY OF SCIENCE</BusinessNameLine1Txt>
      </BusinessName>
      <BusinessNameControlTxt>PUBL</BusinessNameControlTxt>
      <PhoneNum>4156241200</PhoneNum>
      <USAddress>
        <AddressLine1Txt>2112 CHESTNUT ST 310</AddressLine1Txt>
        <CityNm>ALHAMBRA</CityNm>
        <StateAbbreviationCd>CA</StateAbbreviationCd>
        <ZIPCd>91803</ZIPCd>
      </USAddress>
    </Filer>
    <BusinessOfficerGrp>
      <PersonNm>KATHRYN MOTONAGA</PersonNm>
      <PersonTitleTxt>CFO</PersonTitleTxt>
      <PhoneNum>4156241200</PhoneNum>
      <SignatureDt>2025-11-13</SignatureDt>
      <DiscussWithPaidPreparerInd>1</DiscussWithPaidPreparerInd>
    </BusinessOfficerGrp>
    <IRSResponsiblePrtyInfoCurrInd>1</IRSResponsiblePrtyInfoCurrInd>
    <PreparerPersonGrp>
      <PreparerPersonNm>KATY BROWN</PreparerPersonNm>
      <PTIN>P00650274</PTIN>
      <PhoneNum>9257902600</PhoneNum>
      <PreparationDt>2025-11-13</PreparationDt>
    </PreparerPersonGrp>
    <TaxYr>2024</TaxYr>
    <BuildTS>2025-03-06 01:10:19Z</BuildTS>
  </ReturnHeader>
  <ReturnData documentCnt="8">
    <IRS990 documentId="RetDoc1038000001" referenceDocumentId="RetDoc1044400001">
      <AddressChangeInd>X</AddressChangeInd>
      <PrincipalOfficerNm>KATHRYN MOTONAGA</PrincipalOfficerNm>
      <USAddress>
        <AddressLine1Txt>2112 CHESTNUT ST 310</AddressLine1Txt>
        <CityNm>ALHAMBRA</CityNm>
        <StateAbbreviationCd>CA</StateAbbreviationCd>
        <ZIPCd>91803</ZIPCd>
      </USAddress>
      <GrossReceiptsAmt>41547134</GrossReceiptsAmt>
      <GroupReturnForAffiliatesInd>0</GroupReturnForAffiliatesInd>
      <Organization501c3Ind>X</Organization501c3Ind>
      <WebsiteAddressTxt>WWW.PLOS.ORG</WebsiteAddressTxt>
      <TypeOfOrganizationCorpInd>X</TypeOfOrganizationCorpInd>
      <FormationYr>2001</FormationYr>
      <LegalDomicileStateCd>CA</LegalDomicileStateCd>
      <ActivityOrMissionDesc>PLEASE SEE SCHEDULE O FOR COMPLETE DESCRIPTION OF THE ORGANIZATION'S MISSION.</ActivityOrMissionDesc>
      <VotingMembersGoverningBodyCnt>13</VotingMembersGoverningBodyCnt>
      <VotingMembersIndependentCnt>12</VotingMembersIndependentCnt>
      <TotalEmployeeCnt>236</TotalEmployeeCnt>
      <TotalVolunteersCnt>13500</TotalVolunteersCnt>
      <TotalGrossUBIAmt>159134</TotalGrossUBIAmt>
      <NetUnrelatedBusTxblIncmAmt>0</NetUnrelatedBusTxblIncmAmt>
      <PYContributionsGrantsAmt>14248</PYContributionsGrantsAmt>
      <CYContributionsGrantsAmt>2516683</CYContributionsGrantsAmt>
      <PYProgramServiceRevenueAmt>32532613</PYProgramServiceRevenueAmt>
      <CYProgramServiceRevenueAmt>38061535</CYProgramServiceRevenueAmt>
      <PYInvestmentIncomeAmt>484569</PYInvestmentIncomeAmt>
      <CYInvestmentIncomeAmt>757059</CYInvestmentIncomeAmt>
      <PYOtherRevenueAmt>1331</PYOtherRevenueAmt>
      <CYOtherRevenueAmt>126</CYOtherRevenueAmt>
      <PYTotalRevenueAmt>33032761</PYTotalRevenueAmt>
      <CYTotalRevenueAmt>41335403</CYTotalRevenueAmt>
      <PYGrantsAndSimilarPaidAmt>0</PYGrantsAndSimilarPaidAmt>
      <CYGrantsAndSimilarPaidAmt>0</CYGrantsAndSimilarPaidAmt>
      <PYBenefitsPaidToMembersAmt>0</PYBenefitsPaidToMembersAmt>
      <CYBenefitsPaidToMembersAmt>0</CYBenefitsPaidToMembersAmt>
      <PYSalariesCompEmpBnftPaidAmt>21446020</PYSalariesCompEmpBnftPaidAmt>
      <CYSalariesCompEmpBnftPaidAmt>23438680</CYSalariesCompEmpBnftPaidAmt>
      <PYTotalProfFndrsngExpnsAmt>0</PYTotalProfFndrsngExpnsAmt>
      <CYTotalProfFndrsngExpnsAmt>0</CYTotalProfFndrsngExpnsAmt>
      <CYTotalFundraisingExpenseAmt>0</CYTotalFundraisingExpenseAmt>
      <PYOtherExpensesAmt>13484138</PYOtherExpensesAmt>
      <CYOtherExpensesAmt>13546742</CYOtherExpensesAmt>
      <PYTotalExpensesAmt>34930158</PYTotalExpensesAmt>
      <CYTotalExpensesAmt>36985422</CYTotalExpensesAmt>
      <PYRevenuesLessExpensesAmt>-1897397</PYRevenuesLessExpensesAmt>
      <CYRevenuesLessExpensesAmt>4349981</CYRevenuesLessExpensesAmt>
      <TotalAssetsBOYAmt>24736549</TotalAssetsBOYAmt>
      <TotalAssetsEOYAmt>32799863</TotalAssetsEOYAmt>
      <TotalLiabilitiesBOYAmt>6516531</TotalLiabilitiesBOYAmt>
      <TotalLiabilitiesEOYAmt>9775427</TotalLiabilitiesEOYAmt>
      <NetAssetsOrFundBalancesBOYAmt>18220018</NetAssetsOrFundBalancesBOYAmt>
      <NetAssetsOrFundBalancesEOYAmt>23024436</NetAssetsOrFundBalancesEOYAmt>
      <InfoInScheduleOPartIIIInd>X</InfoInScheduleOPartIIIInd>
      <MissionDesc>PLEASE SEE SCHEDULE O FOR COMPLETE DESCRIPTION OF THE ORGANIZATION'S MISSION.</MissionDesc>
      <SignificantNewProgramSrvcInd>0</SignificantNewProgramSrvcInd>
      <SignificantChangeInd>0</SignificantChangeInd>
      <ExpenseAmt>31047002</ExpenseAmt>
      <RevenueAmt>37902401</RevenueAmt>
      <Desc>THE PUBLIC LIBRARY OF SCIENCE (PLOS) ADVANCES ITS CHARITABLE PURPOSE BY DRIVING OPEN SCIENCE FORWARD WITH MEASURABLE, MEANINGFUL CHANGE IN RESEARCH PUBLISHING, POLICY, AND PRACTICE. WE PUBLISH A PORTFOLIO OF PEER-REVIEWED, OPEN ACCESS SCIENTIFIC JOURNALS, REMOVING FINANCIAL, LEGAL, AND TECHNICAL BARRIERS TO ACCESSING RESEARCH BY ENSURING THAT ALL ARTICLES AND ASSOCIATED DATA ARE FREELY AVAILABLE TO THE PUBLIC WITHOUT SUBSCRIPTION FEES. DURING THE TAX YEAR, PLOS PUBLISHED APPROXIMATELY 20,000 PEER-REVIEWED ARTICLES FROM AUTHORS IN 163 COUNTRIES ACROSS 14 SCIENTIFIC JOURNALS AND PROVIDED $5.7 MILLION IN PARTIAL OR FULL ARTICLE PROCESSING CHARGE (APC) WAIVERS TO AUTHORS. 4A. LARGEST PROGRAM SERVICE (BY EXPENSES):SCIENCE AND MEDICAL PUBLISHING ARE THE CORE OF OUR PROGRAM EXPENSE AT OVER $31MILLION. IN 2024 WE PUBLISHED MORE THAN 20,000 RESEARCH ARTICLES DOCUMENTING THE METHODS, OBSERVATIONS AND DISCOVERIES OF A DIVERSE SCIENTIFIC COMMUNITY. THOUSANDS OF RESEARCHERS ACROSS THE GLOBE DEDICATED THEIR TIME AS ACADEMIC EDITORS AND REVIEWERS TO BRINGING THIS WORK TO THE PUBLIC.THE ACTIVITIES INVOLVED IN OUR PUBLISHING OPERATION INCLUDE:- EDITORIAL AND PRODUCTION WORK - RECEIPT AND TRIAGE OF SUBMISSIONS; PERFORMING AND COORDINATING THEIR PEER REVIEW; AND PRODUCTION AND ONLINE PUBLICATION OF ACCEPTED ARTICLES. - MARKETING AND OUTREACH - PROMOTING ARTICLES AND INVITING SUBMISSIONS, INCREASING AWARENESS AND UNDERSTANDING OF OPEN SCIENCE BENEFITS AND PRACTICES, PROMOTING INITIATIVES SUCH AS CALLS FOR PAPERS, AND CREATING RESOURCES FOR AUTHORS AND REVIEWERS.- PUBLISHING ETHICS - HANDLING ETHICAL AND SCIENTIFIC CONCERNS INVOLVING SUBMISSIONS AND PUBLISHED ARTICLES.- ENGAGEMENT WITH RESEARCH COMMUNITIES (E.G. BY ATTENDING AND/OR ORGANIZING CONFERENCES, EVENTS, AND WORKSHOPS) - TO ENSURE THE JOURNALS AND THEIR POLICIES REFLECT THE RESEARCH INTERESTS AND VALUES OF THESE FIELDS AND COMMUNITIES.- MEDIA - COLLABORATING WITH AUTHORS, INSTITUTIONS, SCIENCE MEDIA CENTERS AND JOURNALISTS TO COMMUNICATE RESEARCH TO BOTH SCIENTISTS AND SOCIETY. THE SCOPE OF PLOS JOURNALS PUBLISHING IN 2024 INCLUDES:- PLOS ONE IS A GLOBAL, INCLUSIVE JOURNAL PUBLISHING RESEARCH IN OVER TWO HUNDRED SUBJECT AREAS ACROSS SCIENCE, ENGINEERING, MEDICINE, AND THE RELATED SOCIAL SCIENCES AND HUMANITIES. - PLOS BIOLOGY IS A LEADING LIFE SCIENCE JOURNAL THAT CHAMPIONS HIGH-IMPACT RESEARCH ACROSS ALL DISCIPLINESFROM MOLECULES TO ECOSYSTEMS.- PLOS CLIMATE EMPOWERS GLOBAL, MULTIDISCIPLINARY COLLABORATION, AND ENABLES DECISION-MAKERS AT EVERY LEVEL OF SOCIETY TO TAKE URGENT, EVIDENCE-BASED ACTION ON THE CAUSES AND IMPACTS OF CLIMATE CHANGE.- PLOS COMPLEX SYSTEMS IS A SELECTIVE INTERDISCIPLINARY JOURNAL FOCUSED ON RESEARCH THAT AIDS OUR UNDERSTANDING OF THE WORLD THROUGH A COMPLEXITY LENS, AND FOSTERS DISCUSSION WITHIN THE EMERGING COMPLEXITY COMMUNITY.- PLOS COMPUTATIONAL BIOLOGY IS HOME TO RESEARCH OF SIGNIFICANCE THAT SEEKS TO BETTER UNDERSTAND THE LIVING WORLD THROUGH COMPUTATIONAL METHODS AND CUTTING-EDGE TOOLS.- PLOS DIGITAL HEALTH IS DEDICATED TO ADVANCING EQUITABLE AND DIGITALLY-ENABLED HEALTHCARE, INCREASING TRUST IN NEW TOOLS, AND IMPROVING REPRODUCIBILITY IN THE FIELD OF DIGITAL HEALTH RESEARCH. - PLOS GENETICS PUBLISHES RESEARCH RELATED TO GENETICS AND GENOMICS ACROSS THE FULL BREADTH OF THIS FIELD, FROM MICROBES TO HUMANS AND FROM BASIC SCIENCE TO TRANSLATIONAL APPROACHES.- PLOS GLOBAL PUBLIC HEALTH MAKES WORK OF THE HIGHEST METHODOLOGICAL AND ETHICAL RIGOR OPENLY AVAILABLE TO THE RESEARCHERS, POLICY EXPERTS, AND CLINICIANS WHO DEPEND ON IT TO TACKLE URGENT HEALTH CRISES AND ADVANCE HEALTH EQUITY.- PLOS MEDICINE IS OUR LEADING HEALTH SCIENCE JOURNAL PUBLISHING IMPACTFUL RESEARCH THAT TRANSFORMS GLOBAL HEALTHCARE AND ADVANCES CLINICAL UNDERSTANDING.- PLOS NEGLECTED TROPICAL DISEASES FOCUSES ON RESEARCH ADDRESSING THE NEEDS OF NEGLECTED, FORGOTTEN, AND UNDER-RESOURCED POPULATIONS, WORKING WITH GLOBAL COMMUNITIES AND RESEARCHERS TO IMPROVE THE HEALTH OF ALL.- PLOS PATHOGENS PUBLISHES GROUND-BREAKING RESEARCH ON PATHOGEN BIOLOGY AND HOST-PATHOGEN INTERACTIONS ACROSS ALL LIFE FORMS.- PLOS SUSTAINABILITY AND TRANSFORMATION IS OUR LEADING SUSTAINABILITY SCIENCE JOURNAL BRINGING TOGETHER ORIGINAL RESEARCH, ADDRESSING THE GLOBAL CHALLENGES OF TRANSITIONING TO A MORE SUSTAINABLE AND JUST FUTURE.- PLOS WATER DRIVES CHANGE IN SAFE WATER SANITATION AND HYGIENE, IMPROVING INSIGHTS INTO WATER RESOURCE MANAGEMENT AND IMPACTS ACROSS THE GLOBE.- PLOS MENTAL HEALTH IS DEDICATED TO CONNECTING SCIENTIFIC AND CLINICAL RESEARCH WITH LIVED EXPERIENCE, STRIVING TO ENABLE DEEPER UNDERSTANDING, DISCUSSION, AND ACTION IN ADDRESSING CHALLENGES AND IMPROVING HEALTH AND WELL-BEING AROUND THE WORLD.IN ADDITION TO MANAGING THE ABOVE PORTFOLIO OF JOURNALS, WE ARE COMMITTED TO CATALYZING CHANGE IN SCHOLARLY COMMUNICATION BY FOCUSING ON THREE INTERCONNECTED PILLARS: (1) OPEN SCIENCE PUBLISHING (2) OPEN SCIENCE POLICY AND (3) OPEN SCIENCE PRACTICE. THE FOLLOWING ARE KEY HIGHLIGHTS FROM OUR WORK IN 2024:- OPEN SCIENCE PUBLISHINGWE ARE WORKING TO REDEFINE PUBLISHING AND GO BEYOND THE ARTICLE TO FOSTER RESEARCH COMMUNICATION THAT IS SUSTAINABLE, ACCESSIBLE, AND IMPACTFUL FOR ALL. THIS NEW INITIATIVE, SUPPORTED BY GORDON AND BETTY MOORE FOUNDATION AND ROBERT WOODS JOHNSON FOUNDATION, BUILDS UPON OUR LEGACY OF ADVANCING OPEN SCIENCE OUTPUTS AND REIMAGINING ALTERNATIVE MODELS FOR OPEN ACCESS PUBLISHING. WE ALSO CONTINUED OUR PUBLICATION FEE ASSISTANCE PROGRAMS, DESIGNED TO ENSURE RESEARCHERS WHO LACK FUNDS FOR OPEN ACCESS, CAN STILL MAKE THEIR WORK IMMEDIATELY AND FREELY AVAILABLE WHEN THEY PUBLISH IN OUR JOURNALS. IN 2024 PLOS PROVIDED $5.7 MILLION IN PARTIAL OR FULL ARTICLE PROCESSING CHARGE (APC) WAIVERS TO AUTHORS. - OPEN SCIENCE POLICYWE REGULARLY ENGAGE IN POLICY DISCUSSIONS AS AN ADVOCATE FOR OPEN SCIENCE PRACTICE, A CHAMPION FOR RESEARCHERS, AND A TRUSTED LEADER IN RESEARCH INTEGRITY. IN 2024 WE INTRODUCED SEVERAL NEW POLICIES ACROSS OUR JOURNALS TO STRENGTHEN OUR GUARD AGAINST LARGE-SCALE INTEGRITY THREATS. WE ARE ALSO WORKING IN PARTNERSHIP WITH COALITIONS AND JISC TO ENGAGE A MULTI-STAKEHOLDER WORKING GROUP IN MOVING AWAY FROM ARTICLE-BASED CHARGES. AS PART OF THIS COLLABORATION WE RELEASED A NEW TOOL TO HELP ASSESS EQUITY IN SCHOLARLY COMMUNICATION MODELS- OPEN SCIENCE PRACTICEIN ORDER TO MAKE SCIENTIFIC KNOWLEDGE AS ACCESSIBLE AS POSSIBLE, WE WORK CONSISTENTLY WITH DIVERSE COMMUNITIES TO PROMOTE ADOPTION OF OPEN SCIENCE PRACTICES ACROSS DISCIPLINE, GEOGRAPHY, AND CAREER STAGE. OUR WORK TO ADVANCE OPEN SCIENCE PRACTICE MORE BROADLY IS SUPPORTED BY THE CONTINUED DEVELOPMENT OF OPEN SCIENCE INDICATORS. THIS YEAR WE INTRODUCED A NEW INDICATOR MEASURING STUDY REGISTRATION, CONDUCTED FURTHER RESEARCH TO UNDERSTAND THE NEEDS OF DIVERSE STAKEHOLDERS WITH REGARD TO MEASURING OPEN SCIENCE PRACTICES, AND COLLABORATED WITH THE FRENCH OPEN SCIENCE MONITORING INITIATIVE TO SHARE IDEAS AND ADVOCATE FOR THE FURTHER DEVELOPMENT OF MONITORING SOLUTIONS.</Desc>
      <TotalProgramServiceExpensesAmt>31047002</TotalProgramServiceExpensesAmt>
      <DescribedInSection501c3Ind referenceDocumentId="RetDoc1039100001">1</DescribedInSection501c3Ind>
      <ScheduleBRequiredInd referenceDocumentId="RetDoc1234500001">1</ScheduleBRequiredInd>
      <PoliticalCampaignActyInd>0</PoliticalCampaignActyInd>
      <LobbyingActivitiesInd>0</LobbyingActivitiesInd>
      <SubjectToProxyTaxInd>0</SubjectToProxyTaxInd>
      <DonorAdvisedFundInd referenceDocumentId="RetDoc1040000001">0</DonorAdvisedFundInd>
      <ConservationEasementsInd referenceDocumentId="RetDoc1040000001">0</ConservationEasementsInd>
      <CollectionsOfArtInd referenceDocumentId="RetDoc1040000001">0</CollectionsOfArtInd>
      <CreditCounselingInd referenceDocumentId="RetDoc1040000001">0</CreditCounselingInd>
      <DonorRstrOrQuasiEndowmentsInd>0</DonorRstrOrQuasiEndowmentsInd>
      <ReportLandBuildingEquipmentInd referenceDocumentId="RetDoc1040000001">1</ReportLandBuildingEquipmentInd>
      <ReportInvestmentsOtherSecInd referenceDocumentId="RetDoc1040000001">0</ReportInvestmentsOtherSecInd>
      <ReportProgramRelatedInvstInd referenceDocumentId="RetDoc1040000001">0</ReportProgramRelatedInvstInd>
      <ReportOtherAssetsInd referenceDocumentId="RetDoc1040000001">0</ReportOtherAssetsInd>
      <ReportOtherLiabilitiesInd referenceDocumentId="RetDoc1040000001">0</ReportOtherLiabilitiesInd>
      <IncludeFIN48FootnoteInd referenceDocumentId="RetDoc1040000001">1</IncludeFIN48FootnoteInd>
      <IndependentAuditFinclStmtInd referenceDocumentId="RetDoc1040000001">1</IndependentAuditFinclStmtInd>
      <ConsolidatedAuditFinclStmtInd referenceDocumentId="RetDoc1040000001">0</ConsolidatedAuditFinclStmtInd>
      <SchoolOperatingInd>0</SchoolOperatingInd>
      <ForeignOfficeInd>1</ForeignOfficeInd>
      <ForeignActivitiesInd referenceDocumentId="RetDoc1040500001">1</ForeignActivitiesInd>
      <MoreThan5000KToOrgInd referenceDocumentId="RetDoc1040500001">0</MoreThan5000KToOrgInd>
      <MoreThan5000KToIndividualsInd referenceDocumentId="RetDoc1040500001">0</MoreThan5000KToIndividualsInd>
      <ProfessionalFundraisingInd>0</ProfessionalFundraisingInd>
      <FundraisingActivitiesInd>0</FundraisingActivitiesInd>
      <GamingActivitiesInd>0</GamingActivitiesInd>
      <OperateHospitalInd>0</OperateHospitalInd>
      <GrantsToOrganizationsInd>0</GrantsToOrganizationsInd>
      <GrantsToIndividualsInd>0</GrantsToIndividualsInd>
      <ScheduleJRequiredInd referenceDocumentId="RetDoc1042400001">1</ScheduleJRequiredInd>
      <TaxExemptBondsInd>0</TaxExemptBondsInd>
      <EngagedInExcessBenefitTransInd>0</EngagedInExcessBenefitTransInd>
      <PYExcessBenefitTransInd>0</PYExcessBenefitTransInd>
      <LoanOutstandingInd>0</LoanOutstandingInd>
      <GrantToRelatedPersonInd>0</GrantToRelatedPersonInd>
      <BusinessRlnWithOrgMemInd>0</BusinessRlnWithOrgMemInd>
      <BusinessRlnWithFamMemInd>0</BusinessRlnWithFamMemInd>
      <BusinessRlnWith35CtrlEntInd>0</BusinessRlnWith35CtrlEntInd>
      <DeductibleNonCashContriInd>0</DeductibleNonCashContriInd>
      <DeductibleArtContributionInd>0</DeductibleArtContributionInd>
      <TerminateOperationsInd>0</TerminateOperationsInd>
      <PartialLiquidationInd>0</PartialLiquidationInd>
      <DisregardedEntityInd referenceDocumentId="RetDoc1043500001">0</DisregardedEntityInd>
      <RelatedEntityInd referenceDocumentId="RetDoc1043500001">1</RelatedEntityInd>
      <RelatedOrganizationCtrlEntInd>1</RelatedOrganizationCtrlEntInd>
      <TransactionWithControlEntInd referenceDocumentId="RetDoc1043500001">1</TransactionWithControlEntInd>
      <TrnsfrExmptNonChrtblRltdOrgInd referenceDocumentId="RetDoc1043500001">0</TrnsfrExmptNonChrtblRltdOrgInd>
      <ActivitiesConductedPrtshpInd referenceDocumentId="RetDoc1043500001">0</ActivitiesConductedPrtshpInd>
      <ScheduleORequiredInd>1</ScheduleORequiredInd>
      <IRPDocumentCnt>76</IRPDocumentCnt>
      <IRPDocumentW2GCnt>0</IRPDocumentW2GCnt>
      <BackupWthldComplianceInd>1</BackupWthldComplianceInd>
      <EmployeeCnt>236</EmployeeCnt>
      <EmploymentTaxReturnsFiledInd>1</EmploymentTaxReturnsFiledInd>
      <UnrelatedBusIncmOverLimitInd>1</UnrelatedBusIncmOverLimitInd>
      <Form990TFiledInd>1</Form990TFiledInd>
      <ForeignFinancialAccountInd>1</ForeignFinancialAccountInd>
      <ForeignCountryCd>UK</ForeignCountryCd>
      <ForeignCountryCd>GM</ForeignCountryCd>
      <ProhibitedTaxShelterTransInd>0</ProhibitedTaxShelterTransInd>
      <TaxablePartyNotificationInd>0</TaxablePartyNotificationInd>
      <NondeductibleContributionsInd>0</NondeductibleContributionsInd>
      <QuidProQuoContributionsInd>0</QuidProQuoContributionsInd>
      <Form8282PropertyDisposedOfInd>0</Form8282PropertyDisposedOfInd>
      <RcvFndsToPayPrsnlBnftCntrctInd>0</RcvFndsToPayPrsnlBnftCntrctInd>
      <PayPremiumsPrsnlBnftCntrctInd>0</PayPremiumsPrsnlBnftCntrctInd>
      <IndoorTanningServicesInd>0</IndoorTanningServicesInd>
      <SubjToTaxRmnrtnExPrchtPymtInd>0</SubjToTaxRmnrtnExPrchtPymtInd>
      <SubjectToExcsTaxNetInvstIncInd>0</SubjectToExcsTaxNetInvstIncInd>
      <InfoInScheduleOPartVIInd>X</InfoInScheduleOPartVIInd>
      <GoverningBodyVotingMembersCnt>13</GoverningBodyVotingMembersCnt>
      <IndependentVotingMemberCnt>12</IndependentVotingMemberCnt>
      <FamilyOrBusinessRlnInd>0</FamilyOrBusinessRlnInd>
      <DelegationOfMgmtDutiesInd>0</DelegationOfMgmtDutiesInd>
      <ChangeToOrgDocumentsInd>0</ChangeToOrgDocumentsInd>
      <MaterialDiversionOrMisuseInd>0</MaterialDiversionOrMisuseInd>
      <MembersOrStockholdersInd>0</MembersOrStockholdersInd>
      <ElectionOfBoardMembersInd>0</ElectionOfBoardMembersInd>
      <DecisionsSubjectToApprovaInd>0</DecisionsSubjectToApprovaInd>
      <MinutesOfGoverningBodyInd>1</MinutesOfGoverningBodyInd>
      <MinutesOfCommitteesInd>1</MinutesOfCommitteesInd>
      <OfficerMailingAddressInd>0</OfficerMailingAddressInd>
      <LocalChaptersInd>0</LocalChaptersInd>
      <Form990ProvidedToGvrnBodyInd>1</Form990ProvidedToGvrnBodyInd>
      <ConflictOfInterestPolicyInd>1</ConflictOfInterestPolicyInd>
      <AnnualDisclosureCoveredPrsnInd>1</AnnualDisclosureCoveredPrsnInd>
      <RegularMonitoringEnfrcInd>1</RegularMonitoringEnfrcInd>
      <WhistleblowerPolicyInd>1</WhistleblowerPolicyInd>
      <DocumentRetentionPolicyInd>1</DocumentRetentionPolicyInd>
      <CompensationProcessCEOInd>1</CompensationProcessCEOInd>
      <CompensationProcessOtherInd>1</CompensationProcessOtherInd>
      <InvestmentInJointVentureInd>0</InvestmentInJointVentureInd>
      <StatesWhereCopyOfReturnIsFldCd>CA</StatesWhereCopyOfReturnIsFldCd>
      <OwnWebsiteInd>X</OwnWebsiteInd>
      <UponRequestInd>X</UponRequestInd>
      <BooksInCareOfDetail>
        <BusinessName>
          <BusinessNameLine1Txt>KATHRYN MOTONAGA CHIEF FINANCIAL OFFICER</BusinessNameLine1Txt>
        </BusinessName>
        <PhoneNum>4156241200</PhoneNum>
        <USAddress>
          <AddressLine1Txt>2112 CHESTNUT ST 310</AddressLine1Txt>
          <CityNm>ALHAMBRA</CityNm>
          <StateAbbreviationCd>CA</StateAbbreviationCd>
          <ZIPCd>91803</ZIPCd>
        </USAddress>
      </BooksInCareOfDetail>
      <Form990PartVIISectionAGrp>
        <PersonNm>ALASTAIR ADAM</PersonNm>
        <TitleTxt>BOARD MEMBER</TitleTxt>
        <AverageHoursPerWeekRt>5.00</AverageHoursPerWeekRt>
        <IndividualTrusteeOrDirectorInd>X</IndividualTrusteeOrDirectorInd>
        <ReportableCompFromOrgAmt>0</ReportableCompFromOrgAmt>
        <ReportableCompFromRltdOrgAmt>0</ReportableCompFromRltdOrgAmt>
        <OtherCompensationAmt>0</OtherCompensationAmt>
      </Form990PartVIISectionAGrp>
      <Form990PartVIISectionAGrp>
        <PersonNm>AMANDA ARMOUR</PersonNm>
        <TitleTxt>BOARD MEMBER</TitleTxt>
        <AverageHoursPerWeekRt>5.00</AverageHoursPerWeekRt>
        <IndividualTrusteeOrDirectorInd>X</IndividualTrusteeOrDirectorInd>
        <ReportableCompFromOrgAmt>0</ReportableCompFromOrgAmt>
        <ReportableCompFromRltdOrgAmt>0</ReportableCompFromRltdOrgAmt>
        <OtherCompensationAmt>0</OtherCompensationAmt>
      </Form990PartVIISectionAGrp>
      <Form990PartVIISectionAGrp>
        <PersonNm>SURESH BHAT</PersonNm>
        <TitleTxt>BOARD MEMBER</TitleTxt>
        <AverageHoursPerWeekRt>5.00</AverageHoursPerWeekRt>
        <IndividualTrusteeOrDirectorInd>X</IndividualTrusteeOrDirectorInd>
        <ReportableCompFromOrgAmt>0</ReportableCompFromOrgAmt>
        <ReportableCompFromRltdOrgAmt>0</ReportableCompFromRltdOrgAmt>
        <OtherCompensationAmt>0</OtherCompensationAmt>
      </Form990PartVIISectionAGrp>
      <Form990PartVIISectionAGrp>
        <PersonNm>ISRAEL BOROKINI</PersonNm>
        <TitleTxt>BOARD MEMBER</TitleTxt>
        <AverageHoursPerWeekRt>5.00</AverageHoursPerWeekRt>
        <IndividualTrusteeOrDirectorInd>X</IndividualTrusteeOrDirectorInd>
        <ReportableCompFromOrgAmt>0</ReportableCompFromOrgAmt>
        <ReportableCompFromRltdOrgAmt>0</ReportableCompFromRltdOrgAmt>
        <OtherCompensationAmt>0</OtherCompensationAmt>
      </Form990PartVIISectionAGrp>
      <Form990PartVIISectionAGrp>
        <PersonNm>FERNAN FEDERICI</PersonNm>
        <TitleTxt>BOARD MEMBER</TitleTxt>
        <AverageHoursPerWeekRt>5.00</AverageHoursPerWeekRt>
        <IndividualTrusteeOrDirectorInd>X</IndividualTrusteeOrDirectorInd>
        <ReportableCompFromOrgAmt>0</ReportableCompFromOrgAmt>
        <ReportableCompFromRltdOrgAmt>0</ReportableCompFromRltdOrgAmt>
        <OtherCompensationAmt>0</OtherCompensationAmt>
      </Form990PartVIISectionAGrp>
      <Form990PartVIISectionAGrp>
        <PersonNm>JENNY MACHIDA</PersonNm>
        <TitleTxt>BOARD MEMBER</TitleTxt>
        <AverageHoursPerWeekRt>5.00</AverageHoursPerWeekRt>
        <IndividualTrusteeOrDirectorInd>X</IndividualTrusteeOrDirectorInd>
        <ReportableCompFromOrgAmt>0</ReportableCompFromOrgAmt>
        <ReportableCompFromRltdOrgAmt>0</ReportableCompFromRltdOrgAmt>
        <OtherCompensationAmt>0</OtherCompensationAmt>
      </Form990PartVIISectionAGrp>
      <Form990PartVIISectionAGrp>
        <PersonNm>EMILY SENA</PersonNm>
        <TitleTxt>BOARD MEMBER</TitleTxt>
        <AverageHoursPerWeekRt>5.00</AverageHoursPerWeekRt>
        <IndividualTrusteeOrDirectorInd>X</IndividualTrusteeOrDirectorInd>
        <ReportableCompFromOrgAmt>0</ReportableCompFromOrgAmt>
        <ReportableCompFromRltdOrgAmt>0</ReportableCompFromRltdOrgAmt>
        <OtherCompensationAmt>0</OtherCompensationAmt>
      </Form990PartVIISectionAGrp>
      <Form990PartVIISectionAGrp>
        <PersonNm>STEVEN TOM</PersonNm>
        <TitleTxt>BOARD MEMBER</TitleTxt>
        <AverageHoursPerWeekRt>5.00</AverageHoursPerWeekRt>
        <IndividualTrusteeOrDirectorInd>X</IndividualTrusteeOrDirectorInd>
        <ReportableCompFromOrgAmt>0</ReportableCompFromOrgAmt>
        <ReportableCompFromRltdOrgAmt>0</ReportableCompFromRltdOrgAmt>
        <OtherCompensationAmt>0</OtherCompensationAmt>
      </Form990PartVIISectionAGrp>
      <Form990PartVIISectionAGrp>
        <PersonNm>SIMINE VAZIRE</PersonNm>
        <TitleTxt>BOARD MEMBER</TitleTxt>
        <AverageHoursPerWeekRt>5.00</AverageHoursPerWeekRt>
        <IndividualTrusteeOrDirectorInd>X</IndividualTrusteeOrDirectorInd>
        <ReportableCompFromOrgAmt>0</ReportableCompFromOrgAmt>
        <ReportableCompFromRltdOrgAmt>0</ReportableCompFromRltdOrgAmt>
        <OtherCompensationAmt>0</OtherCompensationAmt>
      </Form990PartVIISectionAGrp>
      <Form990PartVIISectionAGrp>
        <PersonNm>KATHERINE WHITE</PersonNm>
        <TitleTxt>BOARD MEMBER</TitleTxt>
        <AverageHoursPerWeekRt>5.00</AverageHoursPerWeekRt>
        <IndividualTrusteeOrDirectorInd>X</IndividualTrusteeOrDirectorInd>
        <ReportableCompFromOrgAmt>0</ReportableCompFromOrgAmt>
        <ReportableCompFromRltdOrgAmt>0</ReportableCompFromRltdOrgAmt>
        <OtherCompensationAmt>0</OtherCompensationAmt>
      </Form990PartVIISectionAGrp>
      <Form990PartVIISectionAGrp>
        <PersonNm>RICHARD WILDER</PersonNm>
        <TitleTxt>BOARD MEMBER</TitleTxt>
        <AverageHoursPerWeekRt>5.00</AverageHoursPerWeekRt>
        <IndividualTrusteeOrDirectorInd>X</IndividualTrusteeOrDirectorInd>
        <ReportableCompFromOrgAmt>0</ReportableCompFromOrgAmt>
        <ReportableCompFromRltdOrgAmt>0</ReportableCompFromRltdOrgAmt>
        <OtherCompensationAmt>0</OtherCompensationAmt>
      </Form990PartVIISectionAGrp>
      <Form990PartVIISectionAGrp>
        <PersonNm>KEITH YAMAMOTO</PersonNm>
        <TitleTxt>BOARD MEMBER</TitleTxt>
        <AverageHoursPerWeekRt>5.00</AverageHoursPerWeekRt>
        <IndividualTrusteeOrDirectorInd>X</IndividualTrusteeOrDirectorInd>
        <ReportableCompFromOrgAmt>0</ReportableCompFromOrgAmt>
        <ReportableCompFromRltdOrgAmt>0</ReportableCompFromRltdOrgAmt>
        <OtherCompensationAmt>0</OtherCompensationAmt>
      </Form990PartVIISectionAGrp>
      <Form990PartVIISectionAGrp>
        <PersonNm>ALISON MUDDITT</PersonNm>
        <TitleTxt>CHIEF EXECUTIVE OFFICER</TitleTxt>
        <AverageHoursPerWeekRt>40.00</AverageHoursPerWeekRt>
        <IndividualTrusteeOrDirectorInd>X</IndividualTrusteeOrDirectorInd>
        <OfficerInd>X</OfficerInd>
        <ReportableCompFromOrgAmt>502639</ReportableCompFromOrgAmt>
        <ReportableCompFromRltdOrgAmt>0</ReportableCompFromRltdOrgAmt>
        <OtherCompensationAmt>40681</OtherCompensationAmt>
      </Form990PartVIISectionAGrp>
      <Form990PartVIISectionAGrp>
        <PersonNm>KATHRYN MOTONAGA</PersonNm>
        <TitleTxt>CHIEF FINANCIAL OFFICER</TitleTxt>
        <AverageHoursPerWeekRt>40.00</AverageHoursPerWeekRt>
        <OfficerInd>X</OfficerInd>
        <ReportableCompFromOrgAmt>298202</ReportableCompFromOrgAmt>
        <ReportableCompFromRltdOrgAmt>0</ReportableCompFromRltdOrgAmt>
        <OtherCompensationAmt>31911</OtherCompensationAmt>
      </Form990PartVIISectionAGrp>
      <Form990PartVIISectionAGrp>
        <PersonNm>SANDRA UPEGUI</PersonNm>
        <TitleTxt>GEN. COUNSEL &amp; CORPORATE SECRETARY</TitleTxt>
        <AverageHoursPerWeekRt>40.00</AverageHoursPerWeekRt>
        <OfficerInd>X</OfficerInd>
        <ReportableCompFromOrgAmt>209897</ReportableCompFromOrgAmt>
        <ReportableCompFromRltdOrgAmt>0</ReportableCompFromRltdOrgAmt>
        <OtherCompensationAmt>20003</OtherCompensationAmt>
      </Form990PartVIISectionAGrp>
      <Form990PartVIISectionAGrp>
        <PersonNm>REBEKAH DARKSMITH</PersonNm>
        <TitleTxt>CHIEF OF STAFF AND CHIEF MARKETING AND COMMUNICATI</TitleTxt>
        <AverageHoursPerWeekRt>40.00</AverageHoursPerWeekRt>
        <KeyEmployeeInd>X</KeyEmployeeInd>
        <ReportableCompFromOrgAmt>311457</ReportableCompFromOrgAmt>
        <ReportableCompFromRltdOrgAmt>0</ReportableCompFromRltdOrgAmt>
        <OtherCompensationAmt>38465</OtherCompensationAmt>
      </Form990PartVIISectionAGrp>
      <Form990PartVIISectionAGrp>
        <PersonNm>VERONIQUE KIERMER</PersonNm>
        <TitleTxt>PUBLISHER &amp; EXEC EDITOR</TitleTxt>
        <AverageHoursPerWeekRt>40.00</AverageHoursPerWeekRt>
        <KeyEmployeeInd>X</KeyEmployeeInd>
        <ReportableCompFromOrgAmt>314457</ReportableCompFromOrgAmt>
        <ReportableCompFromRltdOrgAmt>0</ReportableCompFromRltdOrgAmt>
        <OtherCompensationAmt>25845</OtherCompensationAmt>
      </Form990PartVIISectionAGrp>
      <Form990PartVIISectionAGrp>
        <PersonNm>JAMES HARNEY</PersonNm>
        <TitleTxt>SENIOR MANAGER, STRATEGIC RESEARCH</TitleTxt>
        <AverageHoursPerWeekRt>40.00</AverageHoursPerWeekRt>
        <HighestCompensatedEmployeeInd>X</HighestCompensatedEmployeeInd>
        <ReportableCompFromOrgAmt>208987</ReportableCompFromOrgAmt>
        <ReportableCompFromRltdOrgAmt>0</ReportableCompFromRltdOrgAmt>
        <OtherCompensationAmt>8530</OtherCompensationAmt>
      </Form990PartVIISectionAGrp>
      <Form990PartVIISectionAGrp>
        <PersonNm>JAYA DOBBYN</PersonNm>
        <TitleTxt>CONTROLLER (THRU 04/24)</TitleTxt>
        <AverageHoursPerWeekRt>40.00</AverageHoursPerWeekRt>
        <HighestCompensatedEmployeeInd>X</HighestCompensatedEmployeeInd>
        <ReportableCompFromOrgAmt>200535</ReportableCompFromOrgAmt>
        <ReportableCompFromRltdOrgAmt>0</ReportableCompFromRltdOrgAmt>
        <OtherCompensationAmt>12567</OtherCompensationAmt>
      </Form990PartVIISectionAGrp>
      <Form990PartVIISectionAGrp>
        <PersonNm>MATTHEW VALDEZ</PersonNm>
        <TitleTxt>PRIN. SW ENGINEER, PLATFORM &amp; ENG</TitleTxt>
        <AverageHoursPerWeekRt>40.00</AverageHoursPerWeekRt>
        <HighestCompensatedEmployeeInd>X</HighestCompensatedEmployeeInd>
        <ReportableCompFromOrgAmt>202081</ReportableCompFromOrgAmt>
        <ReportableCompFromRltdOrgAmt>0</ReportableCompFromRltdOrgAmt>
        <OtherCompensationAmt>29467</OtherCompensationAmt>
      </Form990PartVIISectionAGrp>
      <Form990PartVIISectionAGrp>
        <PersonNm>NIAMH O'CONNOR</PersonNm>
        <TitleTxt>CHIEF PUBLISHING OFFICER</TitleTxt>
        <AverageHoursPerWeekRt>40.00</AverageHoursPerWeekRt>
        <HighestCompensatedEmployeeInd>X</HighestCompensatedEmployeeInd>
        <ReportableCompFromOrgAmt>204320</ReportableCompFromOrgAmt>
        <ReportableCompFromRltdOrgAmt>0</ReportableCompFromRltdOrgAmt>
        <OtherCompensationAmt>6204</OtherCompensationAmt>
      </Form990PartVIISectionAGrp>
      <Form990PartVIISectionAGrp>
        <PersonNm>TOM SCOTT</PersonNm>
        <TitleTxt>CHIEF DIGITAL OFFICER</TitleTxt>
        <AverageHoursPerWeekRt>40.00</AverageHoursPerWeekRt>
        <HighestCompensatedEmployeeInd>X</HighestCompensatedEmployeeInd>
        <ReportableCompFromOrgAmt>203890</ReportableCompFromOrgAmt>
        <ReportableCompFromRltdOrgAmt>0</ReportableCompFromRltdOrgAmt>
        <OtherCompensationAmt>44040</OtherCompensationAmt>
      </Form990PartVIISectionAGrp>
      <Form990PartVIISectionAGrp>
        <PersonNm>AARON DODDS</PersonNm>
        <TitleTxt>EXECUTIVE DIRECTOR RESEARCH AND MGMT(THRU 01/2024)</TitleTxt>
        <AverageHoursPerWeekRt>40.00</AverageHoursPerWeekRt>
        <FormerOfcrDirectorTrusteeInd>X</FormerOfcrDirectorTrusteeInd>
        <ReportableCompFromOrgAmt>101384</ReportableCompFromOrgAmt>
        <ReportableCompFromRltdOrgAmt>0</ReportableCompFromRltdOrgAmt>
        <OtherCompensationAmt>2720</OtherCompensationAmt>
      </Form990PartVIISectionAGrp>
      <TotalReportableCompFromOrgAmt>2757849</TotalReportableCompFromOrgAmt>
      <TotReportableCompRltdOrgAmt>0</TotReportableCompRltdOrgAmt>
      <TotalOtherCompensationAmt>260433</TotalOtherCompensationAmt>
      <IndivRcvdGreaterThan100KCnt>45</IndivRcvdGreaterThan100KCnt>
      <FormerOfcrEmployeesListedInd>1</FormerOfcrEmployeesListedInd>
      <TotalCompGreaterThan150KInd>1</TotalCompGreaterThan150KInd>
      <CompensationFromOtherSrcsInd>0</CompensationFromOtherSrcsInd>
      <ContractorCompensationGrp>
        <ContractorName>
          <BusinessName>
            <BusinessNameLine1Txt>APEX COVANTAGE LLC</BusinessNameLine1Txt>
          </BusinessName>
        </ContractorName>
        <ContractorAddress>
          <USAddress>
            <AddressLine1Txt>4045 SHERIDAN AVENUE SUITE 266</AddressLine1Txt>
            <CityNm>MIAMI BEACH</CityNm>
            <StateAbbreviationCd>FL</StateAbbreviationCd>
            <ZIPCd>33140</ZIPCd>
          </USAddress>
        </ContractorAddress>
        <ServicesDesc>COMPOSITION</ServicesDesc>
        <CompensationAmt>1814542</CompensationAmt>
      </ContractorCompensationGrp>
      <ContractorCompensationGrp>
        <ContractorName>
          <BusinessName>
            <BusinessNameLine1Txt>SPI TECHNOLOGIES INC</BusinessNameLine1Txt>
          </BusinessName>
        </ContractorName>
        <ContractorAddress>
          <ForeignAddress>
            <AddressLine1Txt>SPI BUILDING PASCOR DRIVE</AddressLine1Txt>
            <CityNm>PARANAQUE CITY</CityNm>
            <ProvinceOrStateNm>METRO MANILA</ProvinceOrStateNm>
            <CountryCd>RP</CountryCd>
            <ForeignPostalCd>01700</ForeignPostalCd>
          </ForeignAddress>
        </ContractorAddress>
        <ServicesDesc>TECHNOLOGY SERVICES</ServicesDesc>
        <CompensationAmt>1436796</CompensationAmt>
      </ContractorCompensationGrp>
      <ContractorCompensationGrp>
        <ContractorName>
          <BusinessName>
            <BusinessNameLine1Txt>ARIES SYSTEMS CORPORATION</BusinessNameLine1Txt>
          </BusinessName>
        </ContractorName>
        <ContractorAddress>
          <USAddress>
            <AddressLine1Txt>50 HIGH STREET SUITE 21</AddressLine1Txt>
            <CityNm>NORTH ANDOVER</CityNm>
            <StateAbbreviationCd>MA</StateAbbreviationCd>
            <ZIPCd>01845</ZIPCd>
          </USAddress>
        </ContractorAddress>
        <ServicesDesc>MANUSCRIPT SUBMISSION AND TRACKING SYSTE</ServicesDesc>
        <CompensationAmt>1104930</CompensationAmt>
      </ContractorCompensationGrp>
      <ContractorCompensationGrp>
        <ContractorName>
          <BusinessName>
            <BusinessNameLine1Txt>EDITORIAL OFFICE LTD</BusinessNameLine1Txt>
          </BusinessName>
        </ContractorName>
        <ContractorAddress>
          <USAddress>
            <AddressLine1Txt>8 THE GREEN</AddressLine1Txt>
            <CityNm>DOVER</CityNm>
            <StateAbbreviationCd>DE</StateAbbreviationCd>
            <ZIPCd>19901</ZIPCd>
          </USAddress>
        </ContractorAddress>
        <ServicesDesc>EDITORIAL SERVICES</ServicesDesc>
        <CompensationAmt>505687</CompensationAmt>
      </ContractorCompensationGrp>
      <ContractorCompensationGrp>
        <ContractorName>
          <BusinessName>
            <BusinessNameLine1Txt>CLARIVATE ANALYTICS (US) LLC</BusinessNameLine1Txt>
          </BusinessName>
        </ContractorName>
        <ContractorAddress>
          <USAddress>
            <AddressLine1Txt>3133 WEST FRYE ROAD SUITE 400</AddressLine1Txt>
            <CityNm>CHANDLER</CityNm>
            <StateAbbreviationCd>AZ</StateAbbreviationCd>
            <ZIPCd>85226</ZIPCd>
          </USAddress>
        </ContractorAddress>
        <ServicesDesc>AUTHOR CONNECT SERVICES</ServicesDesc>
        <CompensationAmt>375199</CompensationAmt>
      </ContractorCompensationGrp>
      <CntrctRcvdGreaterThan100KCnt>18</CntrctRcvdGreaterThan100KCnt>
      <AllOtherContributionsAmt>2516683</AllOtherContributionsAmt>
      <TotalContributionsAmt>2516683</TotalContributionsAmt>
      <ProgramServiceRevenueGrp>
        <Desc>PUBLICATION FEES, NET</Desc>
        <BusinessCd>517000</BusinessCd>
        <TotalRevenueColumnAmt>30964133</TotalRevenueColumnAmt>
        <RelatedOrExemptFuncIncomeAmt>30964133</RelatedOrExemptFuncIncomeAmt>
      </ProgramServiceRevenueGrp>
      <ProgramServiceRevenueGrp>
        <Desc>SUBSCRIPTIONS</Desc>
        <BusinessCd>517000</BusinessCd>
        <TotalRevenueColumnAmt>6938268</TotalRevenueColumnAmt>
        <RelatedOrExemptFuncIncomeAmt>6938268</RelatedOrExemptFuncIncomeAmt>
      </ProgramServiceRevenueGrp>
      <ProgramServiceRevenueGrp>
        <Desc>ADVERTISING</Desc>
        <BusinessCd>517000</BusinessCd>
        <TotalRevenueColumnAmt>159134</TotalRevenueColumnAmt>
        <UnrelatedBusinessRevenueAmt>159134</UnrelatedBusinessRevenueAmt>
      </ProgramServiceRevenueGrp>
      <TotalProgramServiceRevenueAmt>38061535</TotalProgramServiceRevenueAmt>
      <InvestmentIncomeGrp>
        <TotalRevenueColumnAmt>792750</TotalRevenueColumnAmt>
        <ExclusionAmt>792750</ExclusionAmt>
      </InvestmentIncomeGrp>
      <RoyaltiesRevenueGrp>
        <TotalRevenueColumnAmt>126</TotalRevenueColumnAmt>
        <ExclusionAmt>126</ExclusionAmt>
      </RoyaltiesRevenueGrp>
      <GrossAmountSalesAssetsGrp>
        <SecuritiesAmt>175976</SecuritiesAmt>
        <OtherAmt>64</OtherAmt>
      </GrossAmountSalesAssetsGrp>
      <LessCostOthBasisSalesExpnssGrp>
        <SecuritiesAmt>179386</SecuritiesAmt>
        <OtherAmt>32345</OtherAmt>
      </LessCostOthBasisSalesExpnssGrp>
      <GainOrLossGrp>
        <SecuritiesAmt>-3410</SecuritiesAmt>
        <OtherAmt>-32281</OtherAmt>
      </GainOrLossGrp>
      <NetGainOrLossInvestmentsGrp>
        <TotalRevenueColumnAmt>-35691</TotalRevenueColumnAmt>
        <ExclusionAmt>-35691</ExclusionAmt>
      </NetGainOrLossInvestmentsGrp>
      <TotalRevenueGrp>
        <TotalRevenueColumnAmt>41335403</TotalRevenueColumnAmt>
        <RelatedOrExemptFuncIncomeAmt>37902401</RelatedOrExemptFuncIncomeAmt>
        <UnrelatedBusinessRevenueAmt>159134</UnrelatedBusinessRevenueAmt>
        <ExclusionAmt>757185</ExclusionAmt>
      </TotalRevenueGrp>
      <CompCurrentOfcrDirectorsGrp>
        <TotalAmt>1793556</TotalAmt>
        <ProgramServicesAmt>340302</ProgramServicesAmt>
        <ManagementAndGeneralAmt>1453254</ManagementAndGeneralAmt>
      </CompCurrentOfcrDirectorsGrp>
      <OtherSalariesAndWagesGrp>
        <TotalAmt>17593900</TotalAmt>
        <ProgramServicesAmt>16038043</ProgramServicesAmt>
        <ManagementAndGeneralAmt>1555857</ManagementAndGeneralAmt>
      </OtherSalariesAndWagesGrp>
      <PensionPlanContributionsGrp>
        <TotalAmt>447009</TotalAmt>
        <ProgramServicesAmt>414948</ProgramServicesAmt>
        <ManagementAndGeneralAmt>32061</ManagementAndGeneralAmt>
      </PensionPlanContributionsGrp>
      <OtherEmployeeBenefitsGrp>
        <TotalAmt>1921408</TotalAmt>
        <ProgramServicesAmt>1706589</ProgramServicesAmt>
        <ManagementAndGeneralAmt>214819</ManagementAndGeneralAmt>
      </OtherEmployeeBenefitsGrp>
      <PayrollTaxesGrp>
        <TotalAmt>1682807</TotalAmt>
        <ProgramServicesAmt>1430958</ProgramServicesAmt>
        <ManagementAndGeneralAmt>251849</ManagementAndGeneralAmt>
      </PayrollTaxesGrp>
      <FeesForServicesLegalGrp>
        <TotalAmt>231440</TotalAmt>
        <ProgramServicesAmt>100288</ProgramServicesAmt>
        <ManagementAndGeneralAmt>131152</ManagementAndGeneralAmt>
      </FeesForServicesLegalGrp>
      <FeesForServicesAccountingGrp>
        <TotalAmt>147430</TotalAmt>
        <ProgramServicesAmt>63884</ProgramServicesAmt>
        <ManagementAndGeneralAmt>83546</ManagementAndGeneralAmt>
      </FeesForServicesAccountingGrp>
      <FeesForSrvcInvstMgmntFeesGrp>
        <TotalAmt>29942</TotalAmt>
        <ManagementAndGeneralAmt>29942</ManagementAndGeneralAmt>
      </FeesForSrvcInvstMgmntFeesGrp>
      <FeesForServicesOtherGrp>
        <TotalAmt>1460712</TotalAmt>
        <ProgramServicesAmt>632957</ProgramServicesAmt>
        <ManagementAndGeneralAmt>827755</ManagementAndGeneralAmt>
      </FeesForServicesOtherGrp>
      <AdvertisingGrp>
        <TotalAmt>301471</TotalAmt>
        <ProgramServicesAmt>301471</ProgramServicesAmt>
      </AdvertisingGrp>
      <OfficeExpensesGrp>
        <TotalAmt>872137</TotalAmt>
        <ProgramServicesAmt>198585</ProgramServicesAmt>
        <ManagementAndGeneralAmt>673552</ManagementAndGeneralAmt>
      </OfficeExpensesGrp>
      <InformationTechnologyGrp>
        <TotalAmt>1965938</TotalAmt>
        <ProgramServicesAmt>1724329</ProgramServicesAmt>
        <ManagementAndGeneralAmt>241609</ManagementAndGeneralAmt>
      </InformationTechnologyGrp>
      <OccupancyGrp>
        <TotalAmt>922239</TotalAmt>
        <ProgramServicesAmt>841512</ProgramServicesAmt>
        <ManagementAndGeneralAmt>80727</ManagementAndGeneralAmt>
      </OccupancyGrp>
      <TravelGrp>
        <TotalAmt>872352</TotalAmt>
        <ProgramServicesAmt>661793</ProgramServicesAmt>
        <ManagementAndGeneralAmt>210559</ManagementAndGeneralAmt>
      </TravelGrp>
      <DepreciationDepletionGrp>
        <TotalAmt>171161</TotalAmt>
        <ProgramServicesAmt>156179</ProgramServicesAmt>
        <ManagementAndGeneralAmt>14982</ManagementAndGeneralAmt>
      </DepreciationDepletionGrp>
      <InsuranceGrp>
        <TotalAmt>105348</TotalAmt>
        <ProgramServicesAmt>96126</ProgramServicesAmt>
        <ManagementAndGeneralAmt>9222</ManagementAndGeneralAmt>
      </InsuranceGrp>
      <OtherExpensesGrp>
        <Desc>PRODUCTION COSTS</Desc>
        <TotalAmt>6236914</TotalAmt>
        <ProgramServicesAmt>6236914</ProgramServicesAmt>
      </OtherExpensesGrp>
      <OtherExpensesGrp>
        <Desc>TRAINING &amp; RECRUITMENT</Desc>
        <TotalAmt>229658</TotalAmt>
        <ProgramServicesAmt>102124</ProgramServicesAmt>
        <ManagementAndGeneralAmt>127534</ManagementAndGeneralAmt>
      </OtherExpensesGrp>
      <TotalFunctionalExpensesGrp>
        <TotalAmt>36985422</TotalAmt>
        <ProgramServicesAmt>31047002</ProgramServicesAmt>
        <ManagementAndGeneralAmt>5938420</ManagementAndGeneralAmt>
        <FundraisingAmt>0</FundraisingAmt>
      </TotalFunctionalExpensesGrp>
      <CashNonInterestBearingGrp>
        <BOYAmt>592276</BOYAmt>
        <EOYAmt>2654549</EOYAmt>
      </CashNonInterestBearingGrp>
      <SavingsAndTempCashInvstGrp>
        <BOYAmt>1465665</BOYAmt>
        <EOYAmt>3477405</EOYAmt>
      </SavingsAndTempCashInvstGrp>
      <PledgesAndGrantsReceivableGrp>
        <BOYAmt>8005</BOYAmt>
        <EOYAmt>760374</EOYAmt>
      </PledgesAndGrantsReceivableGrp>
      <AccountsReceivableGrp>
        <BOYAmt>5588070</BOYAmt>
        <EOYAmt>8131089</EOYAmt>
      </AccountsReceivableGrp>
      <PrepaidExpensesDefrdChargesGrp>
        <BOYAmt>822540</BOYAmt>
        <EOYAmt>1282841</EOYAmt>
      </PrepaidExpensesDefrdChargesGrp>
      <LandBldgEquipCostOrOtherBssAmt>2501900</LandBldgEquipCostOrOtherBssAmt>
      <LandBldgEquipAccumDeprecAmt>2333291</LandBldgEquipAccumDeprecAmt>
      <LandBldgEquipBasisNetGrp>
        <BOYAmt>239760</BOYAmt>
        <EOYAmt>168609</EOYAmt>
      </LandBldgEquipBasisNetGrp>
      <InvestmentsPubTradedSecGrp>
        <BOYAmt>15193088</BOYAmt>
        <EOYAmt>16324996</EOYAmt>
      </InvestmentsPubTradedSecGrp>
      <OtherAssetsTotalGrp>
        <BOYAmt>827145</BOYAmt>
        <EOYAmt>0</EOYAmt>
      </OtherAssetsTotalGrp>
      <TotalAssetsGrp>
        <BOYAmt>24736549</BOYAmt>
        <EOYAmt>32799863</EOYAmt>
      </TotalAssetsGrp>
      <AccountsPayableAccrExpnssGrp>
        <BOYAmt>4075846</BOYAmt>
        <EOYAmt>3967702</EOYAmt>
      </AccountsPayableAccrExpnssGrp>
      <DeferredRevenueGrp>
        <BOYAmt>1523386</BOYAmt>
        <EOYAmt>5807725</EOYAmt>
      </DeferredRevenueGrp>
      <OtherLiabilitiesGrp>
        <BOYAmt>917299</BOYAmt>
        <EOYAmt>0</EOYAmt>
      </OtherLiabilitiesGrp>
      <TotalLiabilitiesGrp>
        <BOYAmt>6516531</BOYAmt>
        <EOYAmt>9775427</EOYAmt>
      </TotalLiabilitiesGrp>
      <OrganizationFollowsFASB117Ind>X</OrganizationFollowsFASB117Ind>
      <NoDonorRestrictionNetAssetsGrp>
        <BOYAmt>18220018</BOYAmt>
        <EOYAmt>20684819</EOYAmt>
      </NoDonorRestrictionNetAssetsGrp>
      <DonorRestrictionNetAssetsGrp>
        <EOYAmt>2339617</EOYAmt>
      </DonorRestrictionNetAssetsGrp>
      <TotalNetAssetsFundBalanceGrp>
        <BOYAmt>18220018</BOYAmt>
        <EOYAmt>23024436</EOYAmt>
      </TotalNetAssetsFundBalanceGrp>
      <TotLiabNetAssetsFundBalanceGrp>
        <BOYAmt>24736549</BOYAmt>
        <EOYAmt>32799863</EOYAmt>
      </TotLiabNetAssetsFundBalanceGrp>
      <ReconcilationRevenueExpnssAmt>4349981</ReconcilationRevenueExpnssAmt>
      <NetUnrlzdGainsLossesInvstAmt>454437</NetUnrlzdGainsLossesInvstAmt>
      <OtherChangesInNetAssetsAmt>0</OtherChangesInNetAssetsAmt>
      <MethodOfAccountingAccrualInd>X</MethodOfAccountingAccrualInd>
      <AccountantCompileOrReviewInd>0</AccountantCompileOrReviewInd>
      <FSAuditedInd>1</FSAuditedInd>
      <FSAuditedBasisGrp>
        <ConsolidatedBasisFinclStmtInd>X</ConsolidatedBasisFinclStmtInd>
      </FSAuditedBasisGrp>
      <AuditCommitteeInd>1</AuditCommitteeInd>
      <FederalGrantAuditRequiredInd>0</FederalGrantAuditRequiredInd>
    </IRS990>
    <IRS990ScheduleA documentId="RetDoc1039100001">
      <PubliclySupportedOrg509a2Ind>X</PubliclySupportedOrg509a2Ind>
      <GiftsGrantsContrisRcvd509Grp>
        <CurrentTaxYearMinus4YearsAmt>60366</CurrentTaxYearMinus4YearsAmt>
        <CurrentTaxYearMinus3YearsAmt>2574373</CurrentTaxYearMinus3YearsAmt>
        <CurrentTaxYearMinus2YearsAmt>43355</CurrentTaxYearMinus2YearsAmt>
        <CurrentTaxYearMinus1YearAmt>14248</CurrentTaxYearMinus1YearAmt>
        <CurrentTaxYearAmt>2516683</CurrentTaxYearAmt>
        <TotalAmt>5209025</TotalAmt>
      </GiftsGrantsContrisRcvd509Grp>
      <GrossReceiptsAdmissionsGrp>
        <CurrentTaxYearMinus4YearsAmt>32485530</CurrentTaxYearMinus4YearsAmt>
        <CurrentTaxYearMinus3YearsAmt>34122386</CurrentTaxYearMinus3YearsAmt>
        <CurrentTaxYearMinus2YearsAmt>34660585</CurrentTaxYearMinus2YearsAmt>
        <CurrentTaxYearMinus1YearAmt>32532613</CurrentTaxYearMinus1YearAmt>
        <CurrentTaxYearAmt>38061535</CurrentTaxYearAmt>
        <TotalAmt>171862649</TotalAmt>
      </GrossReceiptsAdmissionsGrp>
      <Total509Grp>
        <CurrentTaxYearMinus4YearsAmt>32545896</CurrentTaxYearMinus4YearsAmt>
        <CurrentTaxYearMinus3YearsAmt>36696759</CurrentTaxYearMinus3YearsAmt>
        <CurrentTaxYearMinus2YearsAmt>34703940</CurrentTaxYearMinus2YearsAmt>
        <CurrentTaxYearMinus1YearAmt>32546861</CurrentTaxYearMinus1YearAmt>
        <CurrentTaxYearAmt>40578218</CurrentTaxYearAmt>
        <TotalAmt>177071674</TotalAmt>
      </Total509Grp>
      <AmountsRcvdDsqlfyPersonGrp>
        <CurrentTaxYearAmt>2501293</CurrentTaxYearAmt>
        <TotalAmt>2501293</TotalAmt>
      </AmountsRcvdDsqlfyPersonGrp>
      <SubstantialContributorsAmtGrp>
        <TotalAmt>0</TotalAmt>
      </SubstantialContributorsAmtGrp>
      <SubstAndDsqlfyPrsnsTotGrp>
        <CurrentTaxYearAmt>2501293</CurrentTaxYearAmt>
        <TotalAmt>2501293</TotalAmt>
      </SubstAndDsqlfyPrsnsTotGrp>
      <PublicSupportTotal509Amt>174570381</PublicSupportTotal509Amt>
      <GrossInvestmentIncome509Grp>
        <CurrentTaxYearMinus4YearsAmt>415255</CurrentTaxYearMinus4YearsAmt>
        <CurrentTaxYearMinus3YearsAmt>612076</CurrentTaxYearMinus3YearsAmt>
        <CurrentTaxYearMinus2YearsAmt>365743</CurrentTaxYearMinus2YearsAmt>
        <CurrentTaxYearMinus1YearAmt>496702</CurrentTaxYearMinus1YearAmt>
        <CurrentTaxYearAmt>792876</CurrentTaxYearAmt>
        <TotalAmt>2682652</TotalAmt>
      </GrossInvestmentIncome509Grp>
      <InvestmentIncomeAndUBTIGrp>
        <CurrentTaxYearMinus4YearsAmt>415255</CurrentTaxYearMinus4YearsAmt>
        <CurrentTaxYearMinus3YearsAmt>612076</CurrentTaxYearMinus3YearsAmt>
        <CurrentTaxYearMinus2YearsAmt>365743</CurrentTaxYearMinus2YearsAmt>
        <CurrentTaxYearMinus1YearAmt>496702</CurrentTaxYearMinus1YearAmt>
        <CurrentTaxYearAmt>792876</CurrentTaxYearAmt>
        <TotalAmt>2682652</TotalAmt>
      </InvestmentIncomeAndUBTIGrp>
      <OtherIncome509Grp>
        <CurrentTaxYearMinus1YearAmt>1210</CurrentTaxYearMinus1YearAmt>
        <TotalAmt>1210</TotalAmt>
      </OtherIncome509Grp>
      <TotalSupportCalendarYearGrp>
        <CurrentTaxYearMinus4YearsAmt>32961151</CurrentTaxYearMinus4YearsAmt>
        <CurrentTaxYearMinus3YearsAmt>37308835</CurrentTaxYearMinus3YearsAmt>
        <CurrentTaxYearMinus2YearsAmt>35069683</CurrentTaxYearMinus2YearsAmt>
        <CurrentTaxYearMinus1YearAmt>33044773</CurrentTaxYearMinus1YearAmt>
        <CurrentTaxYearAmt>41371094</CurrentTaxYearAmt>
        <TotalAmt>179755536</TotalAmt>
      </TotalSupportCalendarYearGrp>
      <PublicSupportCY509Pct>0.97120</PublicSupportCY509Pct>
      <PublicSupportPY509Pct>0.98630</PublicSupportPY509Pct>
      <InvestmentIncomeCYPct>0.01490</InvestmentIncomeCYPct>
      <InvestmentIncomePYPct>0.01370</InvestmentIncomePYPct>
      <ThirtyThrPctSuprtTestsCY509Ind>X</ThirtyThrPctSuprtTestsCY509Ind>
    </IRS990ScheduleA>
    <IRS990ScheduleB documentId="RetDoc1234500001">
      <ContributorInformationGrp>
        <ContributorNum>RESTRICTED</ContributorNum>
        <ContributorBusinessName>
          <BusinessNameLine1>RESTRICTED</BusinessNameLine1>
        </ContributorBusinessName>
        <ContributorUSAddress>
          <AddressLine1>RESTRICTED</AddressLine1>
          <AddressLine2>RESTRICTED</AddressLine2>
          <City>RESTRICTED</City>
          <State>RESTRICTED</State>
          <ZIPCode>RESTRICTED</ZIPCode>
        </ContributorUSAddress>
        <TotalContributionsAmt>RESTRICTED</TotalContributionsAmt>
      </ContributorInformationGrp>
    </IRS990ScheduleB>
    <IRS990ScheduleD documentId="RetDoc1040000001">
      <EquipmentGrp>
        <OtherCostOrOtherBasisAmt>1554082</OtherCostOrOtherBasisAmt>
        <DepreciationAmt>1385473</DepreciationAmt>
        <BookValueAmt>168609</BookValueAmt>
      </EquipmentGrp>
      <OtherLandBuildingsGrp>
        <OtherCostOrOtherBasisAmt>947818</OtherCostOrOtherBasisAmt>
        <DepreciationAmt>947818</DepreciationAmt>
        <BookValueAmt>0</BookValueAmt>
      </OtherLandBuildingsGrp>
      <TotalBookValueLandBuildingsAmt>168609</TotalBookValueLandBuildingsAmt>
      <FootnoteTextInd>X</FootnoteTextInd>
      <TotalRevEtcAuditedFinclStmtAmt>41759898</TotalRevEtcAuditedFinclStmtAmt>
      <NetUnrealizedGainsInvstAmt>454437</NetUnrealizedGainsInvstAmt>
      <RevenueNotReportedAmt>454437</RevenueNotReportedAmt>
      <RevenueSubtotalAmt>41305461</RevenueSubtotalAmt>
      <InvestmentExpensesNotIncldAmt>29942</InvestmentExpensesNotIncldAmt>
      <RevenueNotReportedFinclStmtAmt>29942</RevenueNotReportedFinclStmtAmt>
      <TotalRevenuePerForm990Amt>41335403</TotalRevenuePerForm990Amt>
      <TotExpnsEtcAuditedFinclStmtAmt>36955480</TotExpnsEtcAuditedFinclStmtAmt>
      <ExpensesNotReportedAmt>0</ExpensesNotReportedAmt>
      <ExpensesSubtotalAmt>36955480</ExpensesSubtotalAmt>
      <InvestmentExpensesNotIncld2Amt>29942</InvestmentExpensesNotIncld2Amt>
      <ExpensesNotRptFinclStmtAmt>29942</ExpensesNotRptFinclStmtAmt>
      <TotalExpensesPerForm990Amt>36985422</TotalExpensesPerForm990Amt>
      <SupplementalInformationDetail>
        <FormAndLineReferenceDesc>PART X, LINE 2:</FormAndLineReferenceDesc>
        <ExplanationTxt>PLOS HAS RECEIVED NOTIFICATION FROM THE INTERNAL REVENUE SERVICE AND THE STATE OF CALIFORNIA THAT IT QUALIFIES FOR TAX-EXEMPT STATUS UNDER SECTION 501(C)(3) OF THE INTERNAL REVENUE CODE AND SECTION 27301D OF THE CALIFORNIA REVENUE AND TAXATION CODE. THE EXEMPTIONS ARE SUBJECT TO PERIODIC REVIEW BY THE FEDERAL AND STATE TAXING AUTHORITIES, AND MANAGEMENT IS CONFIDENT THAT PLOS CONTINUES TO SATISFY ALL FEDERAL AND STATE STATUTES IN ORDER TO QUALIFY FOR CONTINUED TAX EXEMPTION STATUS. PLOS MAY PERIODICALLY RECEIVE UNRELATED BUSINESS INCOME (SUCH AS ADVERTISING INCOME, ETC.) REQUIRING PLOS TO FILE SEPARATE TAX RETURNS UNDER FEDERAL AND STATE STATUTES. PLOS ALSO HAS CERTAIN TRANSACTIONS REQUIRING THE PAYMENT OF ADDITIONAL EMPLOYER TAXES TO HM REVENUE AND CUSTOMS IN THE U.K. UNDER SUCH CONDITIONS, PLOS CALCULATES AND ACCRUES THE APPLICABLE TAXES. PLOS IS NOT SUBJECT TO EXAMINATION BY TAXING AUTHORITIES, NOR WAS FOR ANY PRIOR YEARS. THE GERMAN SUBSIDIARY, PLOS GMBH, IS NOT AN EXEMPT ORGANIZATION AND IS SUBJECT TO EMPLOYER TAXES, CORPORATE INCOME TAX FILINGS, AND PAYMENT OBLIGATIONS TO THE AUTHORITIES IN GERMANY. PLOS CONSULTS WITH AN AGENT, MAZARS GMBH AND CO. KG, IN GERMANY TO TIMELY PREPARE AND FILE THE RETURNS. PLOS ASSESSES TAX POSITIONS TAKEN OR EXPECTED TO BE TAKEN AGAINST MORE-LIKELY-THAN-NOT RECOGNITION THRESHOLD AND MEASUREMENT ATTRIBUTES FOR FINANCIAL STATEMENT RECOGNITION. BASED ON AN ANALYSIS PREPARED BY PLOS, IT WAS DETERMINED THAT PLOS BELIEVES THAT IT HAS APPROPRIATE SUPPORT FOR ANY TAX POSITIONS TAKEN, AND AS SUCH, DOES NOT HAVE ANY UNCERTAIN TAX POSITIONS THAT ARE MATERIAL TO THE FINANCIAL STATEMENTS.</ExplanationTxt>
      </SupplementalInformationDetail>
    </IRS990ScheduleD>
    <IRS990ScheduleF documentId="RetDoc1040500001">
      <AccountActivitiesOutsideUSGrp>
        <RegionTxt>EUROPE (INCLUDING ICELAND &amp; GREENLAND) - ALBANIA, ANDORRA, AUSTRIA, BELGIUM</RegionTxt>
        <OfficesCnt>2</OfficesCnt>
        <EmployeeCnt>110</EmployeeCnt>
        <TypeOfActivitiesConductedTxt>PROGRAM SERVICES</TypeOfActivitiesConductedTxt>
        <SpecificServicesProvidedTxt>EDITORIAL SUPPORT TO SERVE U.S. OPERATIONS.</SpecificServicesProvidedTxt>
        <RegionTotalExpendituresAmt>7143038</RegionTotalExpendituresAmt>
      </AccountActivitiesOutsideUSGrp>
      <AccountActivitiesOutsideUSGrp>
        <RegionTxt>EAST ASIA AND THE PACIFIC - AUSTRALIA, BRUNEI, BURMA, CAMBODIA,</RegionTxt>
        <OfficesCnt>1</OfficesCnt>
        <EmployeeCnt>1</EmployeeCnt>
        <TypeOfActivitiesConductedTxt>PROGRAM SERVICES</TypeOfActivitiesConductedTxt>
        <SpecificServicesProvidedTxt>EDITORIAL SUPPORT TO SERVE U.S. OPERATIONS.</SpecificServicesProvidedTxt>
        <RegionTotalExpendituresAmt>178142</RegionTotalExpendituresAmt>
      </AccountActivitiesOutsideUSGrp>
      <SubtotalOfficesCnt>3</SubtotalOfficesCnt>
      <SubtotalEmployeesCnt>111</SubtotalEmployeesCnt>
      <ContinuationTotalOfficeCnt>0</ContinuationTotalOfficeCnt>
      <ContinuationTotalEmployeeCnt>0</ContinuationTotalEmployeeCnt>
      <TotalOfficeCnt>3</TotalOfficeCnt>
      <TotalEmployeeCnt>111</TotalEmployeeCnt>
      <SubtotalSpentAmt>7321180</SubtotalSpentAmt>
      <ContinuationSpentAmt>0</ContinuationSpentAmt>
      <TotalSpentAmt>7321180</TotalSpentAmt>
      <TransferToForeignCorpInd>0</TransferToForeignCorpInd>
      <InterestInForeignTrustInd>0</InterestInForeignTrustInd>
      <ForeignCorpOwnershipInd>0</ForeignCorpOwnershipInd>
      <PassiveForeignInvestmestCoInd>0</PassiveForeignInvestmestCoInd>
      <ForeignPartnershipInd>0</ForeignPartnershipInd>
      <BoycottCountriesInd>0</BoycottCountriesInd>
    </IRS990ScheduleF>
    <IRS990ScheduleJ documentId="RetDoc1042400001">
      <CompensationCommitteeInd>X</CompensationCommitteeInd>
      <WrittenEmploymentContractInd>X</WrittenEmploymentContractInd>
      <CompensationSurveyInd>X</CompensationSurveyInd>
      <BoardOrCommitteeApprovalInd>X</BoardOrCommitteeApprovalInd>
      <SeverancePaymentInd>1</SeverancePaymentInd>
      <SupplementalNonqualRtrPlanInd>0</SupplementalNonqualRtrPlanInd>
      <EquityBasedCompArrngmInd>0</EquityBasedCompArrngmInd>
      <CompBasedOnRevenueOfFlngOrgInd>0</CompBasedOnRevenueOfFlngOrgInd>
      <CompBsdOnRevRelatedOrgsInd>0</CompBsdOnRevRelatedOrgsInd>
      <CompBsdNetEarnsFlngOrgInd>1</CompBsdNetEarnsFlngOrgInd>
      <CompBsdNetEarnsRltdOrgsInd>0</CompBsdNetEarnsRltdOrgsInd>
      <AnyNonFixedPaymentsInd>0</AnyNonFixedPaymentsInd>
      <InitialContractExceptionInd>0</InitialContractExceptionInd>
      <RltdOrgOfficerTrstKeyEmplGrp>
        <PersonNm>ALISON MUDDITT</PersonNm>
        <TitleTxt>CHIEF EXECUTIVE OFFICER</TitleTxt>
        <BaseCompensationFilingOrgAmt>438639</BaseCompensationFilingOrgAmt>
        <CompensationBasedOnRltdOrgsAmt>0</CompensationBasedOnRltdOrgsAmt>
        <BonusFilingOrganizationAmount>64000</BonusFilingOrganizationAmount>
        <BonusRelatedOrganizationsAmt>0</BonusRelatedOrganizationsAmt>
        <OtherCompensationFilingOrgAmt>0</OtherCompensationFilingOrgAmt>
        <OtherCompensationRltdOrgsAmt>0</OtherCompensationRltdOrgsAmt>
        <DeferredCompensationFlngOrgAmt>13800</DeferredCompensationFlngOrgAmt>
        <DeferredCompRltdOrgsAmt>0</DeferredCompRltdOrgsAmt>
        <NontaxableBenefitsFilingOrgAmt>26881</NontaxableBenefitsFilingOrgAmt>
        <NontaxableBenefitsRltdOrgsAmt>0</NontaxableBenefitsRltdOrgsAmt>
        <TotalCompensationFilingOrgAmt>543320</TotalCompensationFilingOrgAmt>
        <TotalCompensationRltdOrgsAmt>0</TotalCompensationRltdOrgsAmt>
        <CompReportPrior990FilingOrgAmt>0</CompReportPrior990FilingOrgAmt>
        <CompReportPrior990RltdOrgsAmt>0</CompReportPrior990RltdOrgsAmt>
      </RltdOrgOfficerTrstKeyEmplGrp>
      <RltdOrgOfficerTrstKeyEmplGrp>
        <PersonNm>REBEKAH DARKSMITH</PersonNm>
        <TitleTxt>CHIEF OF STAFF AND CHIEF MARKETING A</TitleTxt>
        <BaseCompensationFilingOrgAmt>311457</BaseCompensationFilingOrgAmt>
        <CompensationBasedOnRltdOrgsAmt>0</CompensationBasedOnRltdOrgsAmt>
        <BonusFilingOrganizationAmount>0</BonusFilingOrganizationAmount>
        <BonusRelatedOrganizationsAmt>0</BonusRelatedOrganizationsAmt>
        <OtherCompensationFilingOrgAmt>0</OtherCompensationFilingOrgAmt>
        <OtherCompensationRltdOrgsAmt>0</OtherCompensationRltdOrgsAmt>
        <DeferredCompensationFlngOrgAmt>11584</DeferredCompensationFlngOrgAmt>
        <DeferredCompRltdOrgsAmt>0</DeferredCompRltdOrgsAmt>
        <NontaxableBenefitsFilingOrgAmt>26881</NontaxableBenefitsFilingOrgAmt>
        <NontaxableBenefitsRltdOrgsAmt>0</NontaxableBenefitsRltdOrgsAmt>
        <TotalCompensationFilingOrgAmt>349922</TotalCompensationFilingOrgAmt>
        <TotalCompensationRltdOrgsAmt>0</TotalCompensationRltdOrgsAmt>
        <CompReportPrior990FilingOrgAmt>0</CompReportPrior990FilingOrgAmt>
        <CompReportPrior990RltdOrgsAmt>0</CompReportPrior990RltdOrgsAmt>
      </RltdOrgOfficerTrstKeyEmplGrp>
      <RltdOrgOfficerTrstKeyEmplGrp>
        <PersonNm>VERONIQUE KIERMER</PersonNm>
        <TitleTxt>PUBLISHER &amp; EXEC EDITOR</TitleTxt>
        <BaseCompensationFilingOrgAmt>314457</BaseCompensationFilingOrgAmt>
        <CompensationBasedOnRltdOrgsAmt>0</CompensationBasedOnRltdOrgsAmt>
        <BonusFilingOrganizationAmount>0</BonusFilingOrganizationAmount>
        <BonusRelatedOrganizationsAmt>0</BonusRelatedOrganizationsAmt>
        <OtherCompensationFilingOrgAmt>0</OtherCompensationFilingOrgAmt>
        <OtherCompensationRltdOrgsAmt>0</OtherCompensationRltdOrgsAmt>
        <DeferredCompensationFlngOrgAmt>12559</DeferredCompensationFlngOrgAmt>
        <DeferredCompRltdOrgsAmt>0</DeferredCompRltdOrgsAmt>
        <NontaxableBenefitsFilingOrgAmt>13286</NontaxableBenefitsFilingOrgAmt>
        <NontaxableBenefitsRltdOrgsAmt>0</NontaxableBenefitsRltdOrgsAmt>
        <TotalCompensationFilingOrgAmt>340302</TotalCompensationFilingOrgAmt>
        <TotalCompensationRltdOrgsAmt>0</TotalCompensationRltdOrgsAmt>
        <CompReportPrior990FilingOrgAmt>0</CompReportPrior990FilingOrgAmt>
        <CompReportPrior990RltdOrgsAmt>0</CompReportPrior990RltdOrgsAmt>
      </RltdOrgOfficerTrstKeyEmplGrp>
      <RltdOrgOfficerTrstKeyEmplGrp>
        <PersonNm>KATHRYN MOTONAGA</PersonNm>
        <TitleTxt>CHIEF FINANCIAL OFFICER</TitleTxt>
        <BaseCompensationFilingOrgAmt>298202</BaseCompensationFilingOrgAmt>
        <CompensationBasedOnRltdOrgsAmt>0</CompensationBasedOnRltdOrgsAmt>
        <BonusFilingOrganizationAmount>0</BonusFilingOrganizationAmount>
        <BonusRelatedOrganizationsAmt>0</BonusRelatedOrganizationsAmt>
        <OtherCompensationFilingOrgAmt>0</OtherCompensationFilingOrgAmt>
        <OtherCompensationRltdOrgsAmt>0</OtherCompensationRltdOrgsAmt>
        <DeferredCompensationFlngOrgAmt>10017</DeferredCompensationFlngOrgAmt>
        <DeferredCompRltdOrgsAmt>0</DeferredCompRltdOrgsAmt>
        <NontaxableBenefitsFilingOrgAmt>21894</NontaxableBenefitsFilingOrgAmt>
        <NontaxableBenefitsRltdOrgsAmt>0</NontaxableBenefitsRltdOrgsAmt>
        <TotalCompensationFilingOrgAmt>330113</TotalCompensationFilingOrgAmt>
        <TotalCompensationRltdOrgsAmt>0</TotalCompensationRltdOrgsAmt>
        <CompReportPrior990FilingOrgAmt>0</CompReportPrior990FilingOrgAmt>
        <CompReportPrior990RltdOrgsAmt>0</CompReportPrior990RltdOrgsAmt>
      </RltdOrgOfficerTrstKeyEmplGrp>
      <RltdOrgOfficerTrstKeyEmplGrp>
        <PersonNm>TOM SCOTT</PersonNm>
        <TitleTxt>CHIEF DIGITAL OFFICER</TitleTxt>
        <BaseCompensationFilingOrgAmt>203890</BaseCompensationFilingOrgAmt>
        <CompensationBasedOnRltdOrgsAmt>0</CompensationBasedOnRltdOrgsAmt>
        <BonusFilingOrganizationAmount>0</BonusFilingOrganizationAmount>
        <BonusRelatedOrganizationsAmt>0</BonusRelatedOrganizationsAmt>
        <OtherCompensationFilingOrgAmt>0</OtherCompensationFilingOrgAmt>
        <OtherCompensationRltdOrgsAmt>0</OtherCompensationRltdOrgsAmt>
        <DeferredCompensationFlngOrgAmt>20335</DeferredCompensationFlngOrgAmt>
        <DeferredCompRltdOrgsAmt>0</DeferredCompRltdOrgsAmt>
        <NontaxableBenefitsFilingOrgAmt>23705</NontaxableBenefitsFilingOrgAmt>
        <NontaxableBenefitsRltdOrgsAmt>0</NontaxableBenefitsRltdOrgsAmt>
        <TotalCompensationFilingOrgAmt>247930</TotalCompensationFilingOrgAmt>
        <TotalCompensationRltdOrgsAmt>0</TotalCompensationRltdOrgsAmt>
        <CompReportPrior990FilingOrgAmt>0</CompReportPrior990FilingOrgAmt>
        <CompReportPrior990RltdOrgsAmt>0</CompReportPrior990RltdOrgsAmt>
      </RltdOrgOfficerTrstKeyEmplGrp>
      <RltdOrgOfficerTrstKeyEmplGrp>
        <PersonNm>MATTHEW VALDEZ</PersonNm>
        <TitleTxt>PRIN. SW ENGINEER, PLATFORM &amp; ENG</TitleTxt>
        <BaseCompensationFilingOrgAmt>202081</BaseCompensationFilingOrgAmt>
        <CompensationBasedOnRltdOrgsAmt>0</CompensationBasedOnRltdOrgsAmt>
        <BonusFilingOrganizationAmount>0</BonusFilingOrganizationAmount>
        <BonusRelatedOrganizationsAmt>0</BonusRelatedOrganizationsAmt>
        <OtherCompensationFilingOrgAmt>0</OtherCompensationFilingOrgAmt>
        <OtherCompensationRltdOrgsAmt>0</OtherCompensationRltdOrgsAmt>
        <DeferredCompensationFlngOrgAmt>7573</DeferredCompensationFlngOrgAmt>
        <DeferredCompRltdOrgsAmt>0</DeferredCompRltdOrgsAmt>
        <NontaxableBenefitsFilingOrgAmt>21894</NontaxableBenefitsFilingOrgAmt>
        <NontaxableBenefitsRltdOrgsAmt>0</NontaxableBenefitsRltdOrgsAmt>
        <TotalCompensationFilingOrgAmt>231548</TotalCompensationFilingOrgAmt>
        <TotalCompensationRltdOrgsAmt>0</TotalCompensationRltdOrgsAmt>
        <CompReportPrior990FilingOrgAmt>0</CompReportPrior990FilingOrgAmt>
        <CompReportPrior990RltdOrgsAmt>0</CompReportPrior990RltdOrgsAmt>
      </RltdOrgOfficerTrstKeyEmplGrp>
      <RltdOrgOfficerTrstKeyEmplGrp>
        <PersonNm>SANDRA UPEGUI</PersonNm>
        <TitleTxt>GEN. COUNSEL &amp; CORPORATE SECRETARY</TitleTxt>
        <BaseCompensationFilingOrgAmt>209897</BaseCompensationFilingOrgAmt>
        <CompensationBasedOnRltdOrgsAmt>0</CompensationBasedOnRltdOrgsAmt>
        <BonusFilingOrganizationAmount>0</BonusFilingOrganizationAmount>
        <BonusRelatedOrganizationsAmt>0</BonusRelatedOrganizationsAmt>
        <OtherCompensationFilingOrgAmt>0</OtherCompensationFilingOrgAmt>
        <OtherCompensationRltdOrgsAmt>0</OtherCompensationRltdOrgsAmt>
        <DeferredCompensationFlngOrgAmt>7780</DeferredCompensationFlngOrgAmt>
        <DeferredCompRltdOrgsAmt>0</DeferredCompRltdOrgsAmt>
        <NontaxableBenefitsFilingOrgAmt>12223</NontaxableBenefitsFilingOrgAmt>
        <NontaxableBenefitsRltdOrgsAmt>0</NontaxableBenefitsRltdOrgsAmt>
        <TotalCompensationFilingOrgAmt>229900</TotalCompensationFilingOrgAmt>
        <TotalCompensationRltdOrgsAmt>0</TotalCompensationRltdOrgsAmt>
        <CompReportPrior990FilingOrgAmt>0</CompReportPrior990FilingOrgAmt>
        <CompReportPrior990RltdOrgsAmt>0</CompReportPrior990RltdOrgsAmt>
      </RltdOrgOfficerTrstKeyEmplGrp>
      <RltdOrgOfficerTrstKeyEmplGrp>
        <PersonNm>JAMES HARNEY</PersonNm>
        <TitleTxt>SENIOR MANAGER, STRATEGIC RESEARCH</TitleTxt>
        <BaseCompensationFilingOrgAmt>208987</BaseCompensationFilingOrgAmt>
        <CompensationBasedOnRltdOrgsAmt>0</CompensationBasedOnRltdOrgsAmt>
        <BonusFilingOrganizationAmount>0</BonusFilingOrganizationAmount>
        <BonusRelatedOrganizationsAmt>0</BonusRelatedOrganizationsAmt>
        <OtherCompensationFilingOrgAmt>0</OtherCompensationFilingOrgAmt>
        <OtherCompensationRltdOrgsAmt>0</OtherCompensationRltdOrgsAmt>
        <DeferredCompensationFlngOrgAmt>8345</DeferredCompensationFlngOrgAmt>
        <DeferredCompRltdOrgsAmt>0</DeferredCompRltdOrgsAmt>
        <NontaxableBenefitsFilingOrgAmt>185</NontaxableBenefitsFilingOrgAmt>
        <NontaxableBenefitsRltdOrgsAmt>0</NontaxableBenefitsRltdOrgsAmt>
        <TotalCompensationFilingOrgAmt>217517</TotalCompensationFilingOrgAmt>
        <TotalCompensationRltdOrgsAmt>0</TotalCompensationRltdOrgsAmt>
        <CompReportPrior990FilingOrgAmt>0</CompReportPrior990FilingOrgAmt>
        <CompReportPrior990RltdOrgsAmt>0</CompReportPrior990RltdOrgsAmt>
      </RltdOrgOfficerTrstKeyEmplGrp>
      <RltdOrgOfficerTrstKeyEmplGrp>
        <PersonNm>JAYA DOBBYN</PersonNm>
        <TitleTxt>CONTROLLER (THRU 04/24)</TitleTxt>
        <BaseCompensationFilingOrgAmt>89938</BaseCompensationFilingOrgAmt>
        <CompensationBasedOnRltdOrgsAmt>0</CompensationBasedOnRltdOrgsAmt>
        <BonusFilingOrganizationAmount>0</BonusFilingOrganizationAmount>
        <BonusRelatedOrganizationsAmt>0</BonusRelatedOrganizationsAmt>
        <OtherCompensationFilingOrgAmt>110597</OtherCompensationFilingOrgAmt>
        <OtherCompensationRltdOrgsAmt>0</OtherCompensationRltdOrgsAmt>
        <DeferredCompensationFlngOrgAmt>3607</DeferredCompensationFlngOrgAmt>
        <DeferredCompRltdOrgsAmt>0</DeferredCompRltdOrgsAmt>
        <NontaxableBenefitsFilingOrgAmt>8960</NontaxableBenefitsFilingOrgAmt>
        <NontaxableBenefitsRltdOrgsAmt>0</NontaxableBenefitsRltdOrgsAmt>
        <TotalCompensationFilingOrgAmt>213102</TotalCompensationFilingOrgAmt>
        <TotalCompensationRltdOrgsAmt>0</TotalCompensationRltdOrgsAmt>
        <CompReportPrior990FilingOrgAmt>0</CompReportPrior990FilingOrgAmt>
        <CompReportPrior990RltdOrgsAmt>0</CompReportPrior990RltdOrgsAmt>
      </RltdOrgOfficerTrstKeyEmplGrp>
      <RltdOrgOfficerTrstKeyEmplGrp>
        <PersonNm>NIAMH O'CONNOR</PersonNm>
        <TitleTxt>CHIEF PUBLISHING OFFICER</TitleTxt>
        <BaseCompensationFilingOrgAmt>204320</BaseCompensationFilingOrgAmt>
        <CompensationBasedOnRltdOrgsAmt>0</CompensationBasedOnRltdOrgsAmt>
        <BonusFilingOrganizationAmount>0</BonusFilingOrganizationAmount>
        <BonusRelatedOrganizationsAmt>0</BonusRelatedOrganizationsAmt>
        <OtherCompensationFilingOrgAmt>0</OtherCompensationFilingOrgAmt>
        <OtherCompensationRltdOrgsAmt>0</OtherCompensationRltdOrgsAmt>
        <DeferredCompensationFlngOrgAmt>1099</DeferredCompensationFlngOrgAmt>
        <DeferredCompRltdOrgsAmt>0</DeferredCompRltdOrgsAmt>
        <NontaxableBenefitsFilingOrgAmt>5105</NontaxableBenefitsFilingOrgAmt>
        <NontaxableBenefitsRltdOrgsAmt>0</NontaxableBenefitsRltdOrgsAmt>
        <TotalCompensationFilingOrgAmt>210524</TotalCompensationFilingOrgAmt>
        <TotalCompensationRltdOrgsAmt>0</TotalCompensationRltdOrgsAmt>
        <CompReportPrior990FilingOrgAmt>0</CompReportPrior990FilingOrgAmt>
        <CompReportPrior990RltdOrgsAmt>0</CompReportPrior990RltdOrgsAmt>
      </RltdOrgOfficerTrstKeyEmplGrp>
      <RltdOrgOfficerTrstKeyEmplGrp>
        <PersonNm>AARON DODDS</PersonNm>
        <TitleTxt>EXECUTIVE DIRECTOR RESEARCH AND MGMT</TitleTxt>
        <BaseCompensationFilingOrgAmt>42344</BaseCompensationFilingOrgAmt>
        <CompensationBasedOnRltdOrgsAmt>0</CompensationBasedOnRltdOrgsAmt>
        <BonusFilingOrganizationAmount>0</BonusFilingOrganizationAmount>
        <BonusRelatedOrganizationsAmt>0</BonusRelatedOrganizationsAmt>
        <OtherCompensationFilingOrgAmt>59040</OtherCompensationFilingOrgAmt>
        <OtherCompensationRltdOrgsAmt>0</OtherCompensationRltdOrgsAmt>
        <DeferredCompensationFlngOrgAmt>1701</DeferredCompensationFlngOrgAmt>
        <DeferredCompRltdOrgsAmt>0</DeferredCompRltdOrgsAmt>
        <NontaxableBenefitsFilingOrgAmt>1019</NontaxableBenefitsFilingOrgAmt>
        <NontaxableBenefitsRltdOrgsAmt>0</NontaxableBenefitsRltdOrgsAmt>
        <TotalCompensationFilingOrgAmt>104104</TotalCompensationFilingOrgAmt>
        <TotalCompensationRltdOrgsAmt>0</TotalCompensationRltdOrgsAmt>
        <CompReportPrior990FilingOrgAmt>0</CompReportPrior990FilingOrgAmt>
        <CompReportPrior990RltdOrgsAmt>0</CompReportPrior990RltdOrgsAmt>
      </RltdOrgOfficerTrstKeyEmplGrp>
      <SupplementalInformationDetail>
        <FormAndLineReferenceDesc>PART I, LINE 4A</FormAndLineReferenceDesc>
        <ExplanationTxt>JAYA DOBBYN, FORMER CONTROLLER (THROUGH APRIL 2024), RECEIVED A SEVERANCE PAYMENT OF $110,596.80, WHICH WAS INCLUDED IN HER REPORTED BASE WAGE. AARON DODDS, FORMER EXECUTIVE DIRECTOR OF RESEARCH AND MANAGEMENT (THROUGH JANUARY 2024), RECEIVED A SEVERANCE PAYMENT OF $59,039.88, ALSO INCLUDED IN HIS REPORTED BASE WAGE.</ExplanationTxt>
      </SupplementalInformationDetail>
      <SupplementalInformationDetail>
        <FormAndLineReferenceDesc>PART I, LINE 6</FormAndLineReferenceDesc>
        <ExplanationTxt>IN 2024, THE ORGANIZATION PROVIDED A BONUS PLAN FOR ALL EMPLOYEES THAT WAS CONTINGENT UPON THE ORGANIZATION'S NET EARNINGS. AS PART OF THIS PLAN, THE CEO RECEIVED A NON-FIXED BONUS OF $64,000.</ExplanationTxt>
      </SupplementalInformationDetail>
    </IRS990ScheduleJ>
    <IRS990ScheduleO documentId="RetDoc1044400001">
      <SupplementalInformationDetail>
        <FormAndLineReferenceDesc>PART I, LINE 1</FormAndLineReferenceDesc>
        <ExplanationTxt>PLOS IS A NON-PROFIT ORGANIZATION ON A MISSION TO DRIVE OPEN SCIENCE FORWARD WITH MEASURABLE, MEANINGFUL CHANGE IN RESEARCH PUBLISHING, POLICY, AND PRACTICE. WE BELIEVE IN A BETTER FUTURE WHERE SCIENCE IS OPEN TO ALL, FOR ALL. BUILDING ON A STRONG LEGACY OF PIONEERING INNOVATION, PLOS CONTINUES TO BE A CATALYST IN OPEN SCIENCE, REIMAGINING MODELS TO MEET OPEN SCIENCE PRINCIPLES, REMOVING BARRIERS AND PROMOTING INCLUSION IN KNOWLEDGE CREATION AND SHARING, AND PUBLISHING RESEARCH OUTPUTS THAT ENABLE EVERYONE TO LEARN FROM, REUSE AND BUILD UPON SCIENTIFIC KNOWLEDGE. OUR WORK IS SUPPORTED BY A HIGHLY SKILLED GLOBAL IN-HOUSE TEAM, PARTNERSHIPS WITH LOCAL SCHOLARLY ORGANIZATIONS, AND THE VALUED CONTRIBUTIONS OF A DIVERSE, INTERNATIONAL COMMUNITY OF SCIENTIFIC RESEARCHERS.</ExplanationTxt>
      </SupplementalInformationDetail>
      <SupplementalInformationDetail>
        <FormAndLineReferenceDesc>FORM 990, PART VI, SECTION B, LINE 11B</FormAndLineReferenceDesc>
        <ExplanationTxt>FORM 990 IS PREPARED BY AN OUTSIDE TAX PROFESSIONAL WITH ASSISTANCE FROM THE ORGANIZATION. THE FORM IS THEN REVIEWED BY THE CFO AND MEMBERS FROM THE AUDIT COMMITTEE. AFTER REVIEW AND MODIFICATIONS WHERE NECESSARY, THE FINAL VERSION OF THE TAX RETURN IS PROVIDED TO THE BOARD OF DIRECTORS. THE CFO SIGNS AND FILES ALL REQUIRED TAX FILINGS.</ExplanationTxt>
      </SupplementalInformationDetail>
      <SupplementalInformationDetail>
        <FormAndLineReferenceDesc>FORM 990, PART VI, SECTION B, LINE 12C</FormAndLineReferenceDesc>
        <ExplanationTxt>THE GENERAL COUNSEL AND SECRETARY OVERSEES THE ANNUAL CONFLICT OF INTEREST DISCLOSURE PROCESS. A COMMITTEE OF THE BOARD OF DIRECTORS REVIEWS ALL POTENTIAL CONFLICTS OF INTEREST AT LEAST ANNUALLY. THE CEO AND ALL BOARD MEMBERS ARE REQUIRED TO DISCLOSE (IN WRITING) POTENTIAL CONFLICTS AND ANY RELATED PARTY AFFILIATIONS. LOANS BETWEEN THE ORGANIZATION AND MEMBERS OF MANAGEMENT AND THE BOARD ARE STRICTLY PROHIBITED. THE ORGANIZATION SEEKS FULL TRANSPARENCY ON ALL RELATIONSHIPS. ANY POTENTIAL CONFLICTS (IN FACT OR APPEARANCE) ARE DISCUSSED OPENLY AND RESOLVED IN ACCORDANCE WITH THE ORGANIZATION'S POLICIES AND PROCEDURES.</ExplanationTxt>
      </SupplementalInformationDetail>
      <SupplementalInformationDetail>
        <FormAndLineReferenceDesc>FORM 990, PART VI, SECTION B, LINE 15</FormAndLineReferenceDesc>
        <ExplanationTxt>THE COMPENSATION COMMITTEE OF THE BOARD OF DIRECTORS REVIEWS THE COMPENSATION OF ALL HIGH-LEVEL PERSONNEL ANNUALLY IN ACCORDANCE WITH IRS RULES AND REGULATIONS. EFFORTS ARE MADE TO SECURE COMPENSATION DATA FROM INDUSTRY SOURCES IN ORDER TO DETERMINE COMPETITIVENESS AND APPROPRIATENESS OF SALARIES AND BENEFITS. EVERY EFFORT IS MADE TO ENSURE THAT THE PROCESS IS THOROUGH AND TRANSPARENT IN ACCORDANCE WITH APPROPRIATE GOVERNANCE AND REGULATORY COMPLIANCE. THIS IS PERFORMED ANNUALLY AND THE PROCESS IS MANAGED BY THE HR DIRECTOR. REVIEW ENCOMPASSES CEO, CFO, EXECUTIVES AND KEY EMPLOYEES.</ExplanationTxt>
      </SupplementalInformationDetail>
      <SupplementalInformationDetail>
        <FormAndLineReferenceDesc>FORM 990, PART VI, SECTION C, LINE 19</FormAndLineReferenceDesc>
        <ExplanationTxt>PLOS MAKES ITS GOVERNING DOCUMENTS, CONFLICT OF INTEREST POLICY, AND FINANCIAL STATEMENTS AVAILABLE TO THE PUBLIC UPON REQUEST. THESE DOCUMENTS ARE ALSO POSTED ON OUR WEBSITE.</ExplanationTxt>
      </SupplementalInformationDetail>
    </IRS990ScheduleO>
    <IRS990ScheduleR documentId="RetDoc1043500001">
      <IdRelatedOrgTxblCorpTrGrp>
        <RelatedOrganizationName>
          <BusinessNameLine1Txt>PLOS GMBH</BusinessNameLine1Txt>
        </RelatedOrganizationName>
        <ForeignAddress>
          <AddressLine1Txt>FRIEDRICHSTR 155</AddressLine1Txt>
          <CityNm>BERLIN</CityNm>
          <ProvinceOrStateNm>BRANDENBURG</ProvinceOrStateNm>
          <CountryCd>GM</CountryCd>
          <ForeignPostalCd>10117</ForeignPostalCd>
        </ForeignAddress>
        <PrimaryActivitiesTxt>RESEARCH</PrimaryActivitiesTxt>
        <LegalDomicileForeignCountryCd>GM</LegalDomicileForeignCountryCd>
        <DirectControllingEntityName>
          <BusinessNameLine1Txt>PUBLIC LIBRARY OF SCIENCE</BusinessNameLine1Txt>
        </DirectControllingEntityName>
        <EntityTypeTxt>C</EntityTypeTxt>
        <ShareOfTotalIncomeAmt>33335</ShareOfTotalIncomeAmt>
        <ShareOfEOYAssetsAmt>306751</ShareOfEOYAssetsAmt>
        <OwnershipPct>1.00000</OwnershipPct>
        <ControlledOrganizationInd>1</ControlledOrganizationInd>
      </IdRelatedOrgTxblCorpTrGrp>
      <IdRelatedOrgTxblCorpTrGrp>
        <RelatedOrganizationName>
          <BusinessNameLine1Txt>PLOS PTE</BusinessNameLine1Txt>
        </RelatedOrganizationName>
        <ForeignAddress>
          <AddressLine1Txt>135 CECIL STREET 10-01 PHILIPPIN</AddressLine1Txt>
          <CityNm>SINGAPORE</CityNm>
          <CountryCd>SN</CountryCd>
          <ForeignPostalCd>69536</ForeignPostalCd>
        </ForeignAddress>
        <PrimaryActivitiesTxt>RESEARCH</PrimaryActivitiesTxt>
        <LegalDomicileForeignCountryCd>SN</LegalDomicileForeignCountryCd>
        <DirectControllingEntityName>
          <BusinessNameLine1Txt>PUBLIC LIBRARY OF SCIENCE</BusinessNameLine1Txt>
        </DirectControllingEntityName>
        <EntityTypeTxt>C</EntityTypeTxt>
        <ShareOfTotalIncomeAmt>-3028</ShareOfTotalIncomeAmt>
        <ShareOfEOYAssetsAmt>22652</ShareOfEOYAssetsAmt>
        <OwnershipPct>1.00000</OwnershipPct>
        <ControlledOrganizationInd>1</ControlledOrganizationInd>
      </IdRelatedOrgTxblCorpTrGrp>
      <ReceiptOfIntAnntsRntsRyltsInd>0</ReceiptOfIntAnntsRntsRyltsInd>
      <GiftGrntOrCapContriToOthOrgInd>0</GiftGrntOrCapContriToOthOrgInd>
      <GiftGrntCapContriFromOthOrgInd>0</GiftGrntCapContriFromOthOrgInd>
      <LoansOrGuaranteesToOtherOrgInd>0</LoansOrGuaranteesToOtherOrgInd>
      <LoansOrGuaranteesFromOthOrgInd>0</LoansOrGuaranteesFromOthOrgInd>
      <DivRelatedOrganizationInd>0</DivRelatedOrganizationInd>
      <AssetSaleToOtherOrgInd>0</AssetSaleToOtherOrgInd>
      <AssetPurchaseFromOtherOrgInd>0</AssetPurchaseFromOtherOrgInd>
      <AssetExchangeInd>0</AssetExchangeInd>
      <RentalOfFacilitiesToOthOrgInd>0</RentalOfFacilitiesToOthOrgInd>
      <RentalOfFcltsFromOthOrgInd>0</RentalOfFcltsFromOthOrgInd>
      <PerformOfServicesForOthOrgInd>1</PerformOfServicesForOthOrgInd>
      <PerformOfServicesByOtherOrgInd>0</PerformOfServicesByOtherOrgInd>
      <SharingOfFacilitiesInd>0</SharingOfFacilitiesInd>
      <PaidEmployeesSharingInd>0</PaidEmployeesSharingInd>
      <ReimbursementPaidToOtherOrgInd>0</ReimbursementPaidToOtherOrgInd>
      <ReimbursementPaidByOtherOrgInd>0</ReimbursementPaidByOtherOrgInd>
      <TransferToOtherOrgInd>0</TransferToOtherOrgInd>
      <TransferFromOtherOrgInd>0</TransferFromOtherOrgInd>
      <TransactionsRelatedOrgGrp>
        <OtherOrganizationName>
          <BusinessNameLine1Txt>PLOS GMBH</BusinessNameLine1Txt>
        </OtherOrganizationName>
        <TransactionTypeTxt>L</TransactionTypeTxt>
        <InvolvedAmt>850525</InvolvedAmt>
        <MethodOfAmountDeterminationTxt>SERVICE AGREEMENT</MethodOfAmountDeterminationTxt>
      </TransactionsRelatedOrgGrp>
      <TransactionsRelatedOrgGrp>
        <OtherOrganizationName>
          <BusinessNameLine1Txt>PLOS PTE</BusinessNameLine1Txt>
        </OtherOrganizationName>
        <TransactionTypeTxt>L</TransactionTypeTxt>
        <InvolvedAmt>257626</InvolvedAmt>
        <MethodOfAmountDeterminationTxt>SERVICE AGREEMENT</MethodOfAmountDeterminationTxt>
      </TransactionsRelatedOrgGrp>
    </IRS990ScheduleR>
  </ReturnData>
</Return>