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        <FormAndLineReferenceDesc>PART I, LINE 4A</FormAndLineReferenceDesc>
        <ExplanationTxt>CLARK HARTSOCK RECEIVED SEVERANCE PAYMENT OF $29,356 RELATING TO TERMINATED EMPLOYMENT ENDING DECEMBER 7, 2018. LOUISE PAGE RECEIVED SEVERANCE PAYMENT OF $66,016 RELATING TO TERMINATED EMPLOYMENT ENDING OCTOBER 31, 2018. CATHERINE RAYHILL RECEIVED SEVERANCE PAYMENT OF $88,512 RELATING TO TERMINATED EMPLOYMENT ENDING APRIL 13, 2018.</ExplanationTxt>
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        <FormAndLineReferenceDesc>PART I, LINE 1</FormAndLineReferenceDesc>
        <ExplanationTxt>PLOS (PUBLIC LIBRARY OF SCIENCE) IS A LEADING NONPROFIT OPEN ACCESS (OA) PUBLISHER DEDICATED TO ACCELERATING PROGRESS IN SCIENCE AND MEDICINE BY LEADING A TRANSFORMATION IN HOW RESEARCHERS COMMUNICATE WITH ONE ANOTHER AND THE PUBLIC. THE PLOS SUITE OF JOURNALS CONTAINS RIGOROUSLY PEER-REVIEWED OA RESEARCH ARTICLES FROM ALL AREAS OF SCIENCE AND MEDICINE, TOGETHER WITH EXPERT COMMENTARY AND ANALYSIS. PLOS BELIEVES SCIENTIFIC IDEAS AND DISCOVERIES ARE A PUBLIC GOOD AND SUPPORTS OA TO MAKE SCIENTIFIC ARTICLES IMMEDIATELY AND FREELY AVAILABLE WITHOUT CHARGE OR OTHER BARRIERS. WE SEEK TO OPTIMIZE THE OPENNESS AND INTEGRITY OF THE PUBLICATION PROCESS TO ENSURE THAT RESEARCH OUTCOMES ARE DISCOVERABLE, ACCESSIBLE AND AVAILABLE FOR DISCUSSION AND THAT SCIENCE COMMUNICATION IS CONSTRUCTIVE, TRANSPARENT AND VERIFIABLE. WE STRIVE TO IMPLEMENT POLICIES AND INNOVATIONS THAT PROMOTE REPRODUCIBILITY, CREDIT AND ACCOUNTABILITY IN ORDER TO FOSTER A CULTURE OF OPEN SCIENCE, WITH OPEN DATA, EARLY SHARING OF WORK AND CLEAR CONTRIBUTOR RECOGNITION. OUR WORK IN THESE AREAS IS SUPPORTED BY THE EFFORTS OF A HIGHLY DIVERSE, INTERNATIONAL COMMUNITY OF SCIENTIFIC RESEARCHERS FROM HUNDREDS OF RESEARCH AREAS WHO PUBLISH AND REVIEW RIGOROUS RESEARCH OF ALL TYPES, ENSURING THAT IT IS MADE ACCESSIBLE TO THE WORLD. SINCE 2003, PLOS HAS PUBLISHED MORE THAN 236,000 ARTICLES. IN 2018 ALONE PLOS PUBLISHED SLIGHTLY MORE THAN 20,000 RESEARCH ARTICLES DOCUMENTING THE OBSERVATIONS AND DISCOVERIES OF A DIVERSE SCIENTIFIC COMMUNITY. A GLOBAL POOL OF MORE THAN 11,000 ACADEMIC EDITORS AND 65,000 REVIEWERS CONTRIBUTED TO BRINGING THIS WORK TO THE PUBLIC.</ExplanationTxt>
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        <FormAndLineReferenceDesc>FORM 990, PART VI, SECTION B, LINE 11B</FormAndLineReferenceDesc>
        <ExplanationTxt>FORM 990 IS PREPARED BY AN OUTSIDE TAX PROFESSIONAL WITH ASSISTANCE FROM THE ORGANIZATION. THE FORM IS THEN REVIEWED BY THE CFO AND MEMBERS FROM THE AUDIT COMMITTEE. AFTER REVIEW AND MODIFICATIONS WHERE NECESSARY, THE FINAL VERSION OF THE TAX RETURN IS PROVIDED TO THE BOARD OF DIRECTORS. THE CFO SIGNS AND FILES ALL REQUIRED TAX FILINGS.</ExplanationTxt>
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        <FormAndLineReferenceDesc>FORM 990, PART VI, SECTION B, LINE 12C</FormAndLineReferenceDesc>
        <ExplanationTxt>THE GENERAL COUNSEL AND SECRETARY OVERSEES THE ANNUAL CONFLICT OF INTEREST DISCLOSURE PROCESS. A COMMITTEE OF THE BOARD OF DIRECTORS REVIEWS ALL POTENTIAL CONFLICTS OF INTEREST AT LEAST ANNUALLY. THE CEO AND ALL BOARD MEMBERS ARE REQUIRED TO DISCLOSE (IN WRITING) POTENTIAL CONFLICTS AND ANY RELATED PARTY AFFILIATIONS. LOANS BETWEEN THE ORGANIZATION AND MEMBERS OF MANAGEMENT AND THE BOARD ARE STRICTLY PROHIBITED. THE ORGANIZATION SEEKS FULL TRANSPARENCY ON ALL RELATIONSHIPS. ANY POTENTIAL CONFLICTS (IN FACT OR APPEARANCE) ARE DISCUSSED OPENLY AND RESOLVED IN ACCORDANCE WITH THE ORGANIZATION'S POLICIES AND PROCEDURES.</ExplanationTxt>
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        <FormAndLineReferenceDesc>FORM 990, PART VI, SECTION B, LINE 15</FormAndLineReferenceDesc>
        <ExplanationTxt>THE COMPENSATION COMMITTEE OF THE BOARD OF DIRECTORS REVIEWS THE COMPENSATION OF ALL HIGH-LEVEL PERSONNEL ANNUALLY IN ACCORDANCE WITH IRS RULES AND REGULATIONS. EFFORTS ARE MADE TO SECURE COMPENSATION DATA FROM INDUSTRY SOURCES IN ORDER TO DETERMINE COMPETITIVENESS AND APPROPRIATENESS OF SALARIES AND BENEFITS. EVERY EFFORT IS MADE TO ENSURE THAT THE PROCESS IS THOROUGH AND TRANSPARENT IN ACCORDANCE WITH APPROPRIATE GOVERNANCE AND REGULATORY COMPLIANCE. THIS IS PERFORMED ANNUALLY AND THE PROCESS IS MANAGED BY THE HR DIRECTOR. REVIEW ENCOMPASSES CEO, CFO, EXECUTIVES AND KEY EMPLOYEES.</ExplanationTxt>
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        <FormAndLineReferenceDesc>FORM 990, PART VI, SECTION C, LINE 19</FormAndLineReferenceDesc>
        <ExplanationTxt>PLOS MAKES ITS GOVERNING DOCUMENTS, CONFLICT OF INTEREST POLICY, AND FINANCIAL STATEMENTS AVAILABLE TO THE PUBLIC. THESE DOCUMENTS ARE ALSO POSTED ON OUR WEBSITE.</ExplanationTxt>
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        <FormAndLineReferenceDesc>FORM 990, PART XI, LINE 9:</FormAndLineReferenceDesc>
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