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        <PersonNm>TODOR I GRIGOROV</PersonNm>
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        <PersonNm>JAQUELINE J MORALES</PersonNm>
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        <PersonNm>VERONIQUE KIERMER</PersonNm>
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        <PersonNm>REBEKAH DARKSMITH</PersonNm>
        <TitleTxt>CHIEF MARKETING OFFICER</TitleTxt>
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      <Form990PartVIISectionAGrp>
        <PersonNm>ANN MICHAEL</PersonNm>
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        <PersonNm>KRISTINA MARTIN</PersonNm>
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        <PersonNm>NIAMH O'CONNOR</PersonNm>
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      <Form990PartVIISectionAGrp>
        <PersonNm>AARON DODDS</PersonNm>
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        <PersonNm>CLARE E DEAN</PersonNm>
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        <PersonNm>JOERG D HEBER</PersonNm>
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      <Form990PartVIISectionAGrp>
        <PersonNm>CHRISTIAN A HAUMESSER</PersonNm>
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      <Form990PartVIISectionAGrp>
        <PersonNm>MIDORI BAER</PersonNm>
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            <BusinessNameLine1Txt>APEX COVANTAGE LLC</BusinessNameLine1Txt>
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          <USAddress>
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      <ContractorCompensationGrp>
        <ContractorName>
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        <ContractorAddress>
          <ForeignAddress>
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      <ContractorCompensationGrp>
        <ContractorName>
          <BusinessName>
            <BusinessNameLine1Txt>J&amp;J EDITORIAL</BusinessNameLine1Txt>
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      <ContractorCompensationGrp>
        <ContractorName>
          <BusinessName>
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          <USAddress>
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      <ContractorCompensationGrp>
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          <BusinessName>
            <BusinessNameLine1Txt>CLARIVATE ANALYTICS (US) LLC</BusinessNameLine1Txt>
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          <USAddress>
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        </ContractorAddress>
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      <TotalContributionsAmt>60366</TotalContributionsAmt>
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        <BusinessCd>519130</BusinessCd>
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      <ProgramServiceRevenueGrp>
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        <BusinessCd>519130</BusinessCd>
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      <ProgramServiceRevenueGrp>
        <Desc>REPRINTS</Desc>
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        <FormAndLineReferenceDesc>PART I, LINE 7</FormAndLineReferenceDesc>
        <ExplanationTxt>THE ORGANIZATION'S CEO (ALISON MUDDITT) RECEIVES NON-FIXED BONUS PAYMENTS. THESE AMOUNTS ARE BASED ON BOARD APPROVAL.</ExplanationTxt>
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      <SupplementalInformationDetail>
        <FormAndLineReferenceDesc>PART I, LINE 1</FormAndLineReferenceDesc>
        <ExplanationTxt>PLOS (PUBLIC LIBRARY OF SCIENCE) IS A NONPROFIT, OPEN ACCESS PUBLISHER EMPOWERING RESEARCHERS TO ACCELERATE PROGRESS IN SCIENCE AND MEDICINE BY LEADING A TRANSFORMATION IN RESEARCH COMMUNICATION. WE HAVE BEEN BREAKING BOUNDARIES SINCE OUR FOUNDING IN 2001. PLOS JOURNALS PROPELLED THE MOVEMENT FOR OA ALTERNATIVES TO SUBSCRIPTION JOURNALS. WE ESTABLISHED THE FIRST MULTI-DISCIPLINARY PUBLICATION INCLUSIVE OF ALL EXCELLENT RESEARCH REGARDLESS OF NOVELTY OR IMPACT AND DEMONSTRATED THE IMPORTANCE OF OPEN DATA AVAILABILITY. AS OPEN SCIENCE ADVANCES, WE CONTINUE TO EXPERIMENT TO PROVIDE MORE OPPORTUNITIES, CHOICE, AND CONTEXT FOR READERS AND RESEARCHERS. PLOS PUBLISHES A SUITE OF INFLUENTIAL OPEN ACCESS JOURNALS ACROSS ALL AREAS OF SCIENCE AND MEDICINE. RIGOROUSLY REPORTED, PEER REVIEWED, AND IMMEDIATELY AVAILABLE WITHOUT RESTRICTIONS, THE JOURNALS PROMOTE THE WIDEST READERSHIP AND IMPACT POSSIBLE. WE STRIVE TO IMPLEMENT POLICIES AND INNOVATIONS THAT PROMOTE REPRODUCIBILITY, CREDIT, AND ACCOUNTABILITY TO FOSTER A CULTURE OF OPEN SCIENCE. OUR WORK IN THESE AREAS IS SUPPORTED BY THE EFFORTS OF A DIVERSE, INTERNATIONAL COMMUNITY OF SCIENTIFIC RESEARCHERS FROM HUNDREDS OF RESEARCH AREAS WHO PUBLISH AND REVIEW RIGOROUS RESEARCH OF ALL TYPES. SINCE 2003, PLOS HAS PUBLISHED MORE THAN 260,000 RESEARCH ARTICLES. IN 2020 ALONE PLOS PUBLISHED MORE THAN 19,000 RESEARCH ARTICLES DOCUMENTING THE OBSERVATIONS AND DISCOVERIES OF A DIVERSE SCIENTIFIC COMMUNITY. THOUSANDS OF RESEARCHERS ACROSS THE GLOBE DEDICATED THEIR TIME AS ACADEMIC EDITORS AND REVIEWERS TO BRINGING THIS WORK TO THE PUBLIC.</ExplanationTxt>
      </SupplementalInformationDetail>
      <SupplementalInformationDetail>
        <FormAndLineReferenceDesc>FORM 990, PART VI, SECTION B, LINE 11B</FormAndLineReferenceDesc>
        <ExplanationTxt>FORM 990 IS PREPARED BY AN OUTSIDE TAX PROFESSIONAL WITH ASSISTANCE FROM THE ORGANIZATION. THE FORM IS THEN REVIEWED BY THE CFO AND MEMBERS FROM THE AUDIT COMMITTEE. AFTER REVIEW AND MODIFICATIONS WHERE NECESSARY, THE FINAL VERSION OF THE TAX RETURN IS PROVIDED TO THE BOARD OF DIRECTORS. THE CFO SIGNS AND FILES ALL REQUIRED TAX FILINGS.</ExplanationTxt>
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      <SupplementalInformationDetail>
        <FormAndLineReferenceDesc>FORM 990, PART VI, SECTION B, LINE 12C</FormAndLineReferenceDesc>
        <ExplanationTxt>THE GENERAL COUNSEL AND SECRETARY OVERSEES THE ANNUAL CONFLICT OF INTEREST DISCLOSURE PROCESS. A COMMITTEE OF THE BOARD OF DIRECTORS REVIEWS ALL POTENTIAL CONFLICTS OF INTEREST AT LEAST ANNUALLY. THE CEO AND ALL BOARD MEMBERS ARE REQUIRED TO DISCLOSE (IN WRITING) POTENTIAL CONFLICTS AND ANY RELATED PARTY AFFILIATIONS. LOANS BETWEEN THE ORGANIZATION AND MEMBERS OF MANAGEMENT AND THE BOARD ARE STRICTLY PROHIBITED. THE ORGANIZATION SEEKS FULL TRANSPARENCY ON ALL RELATIONSHIPS. ANY POTENTIAL CONFLICTS (IN FACT OR APPEARANCE) ARE DISCUSSED OPENLY AND RESOLVED IN ACCORDANCE WITH THE ORGANIZATION'S POLICIES AND PROCEDURES.</ExplanationTxt>
      </SupplementalInformationDetail>
      <SupplementalInformationDetail>
        <FormAndLineReferenceDesc>FORM 990, PART VI, SECTION B, LINE 15</FormAndLineReferenceDesc>
        <ExplanationTxt>THE COMPENSATION COMMITTEE OF THE BOARD OF DIRECTORS REVIEWS THE COMPENSATION OF ALL HIGH-LEVEL PERSONNEL ANNUALLY IN ACCORDANCE WITH IRS RULES AND REGULATIONS. EFFORTS ARE MADE TO SECURE COMPENSATION DATA FROM INDUSTRY SOURCES IN ORDER TO DETERMINE COMPETITIVENESS AND APPROPRIATENESS OF SALARIES AND BENEFITS. EVERY EFFORT IS MADE TO ENSURE THAT THE PROCESS IS THOROUGH AND TRANSPARENT IN ACCORDANCE WITH APPROPRIATE GOVERNANCE AND REGULATORY COMPLIANCE. THIS IS PERFORMED ANNUALLY AND THE PROCESS IS MANAGED BY THE HR DIRECTOR. REVIEW ENCOMPASSES CEO, CFO, EXECUTIVES AND KEY EMPLOYEES.</ExplanationTxt>
      </SupplementalInformationDetail>
      <SupplementalInformationDetail>
        <FormAndLineReferenceDesc>FORM 990, PART VI, SECTION C, LINE 19</FormAndLineReferenceDesc>
        <ExplanationTxt>PLOS MAKES ITS GOVERNING DOCUMENTS, CONFLICT OF INTEREST POLICY, AND FINANCIAL STATEMENTS AVAILABLE TO THE PUBLIC UPON REQUEST. THESE DOCUMENTS ARE ALSO POSTED ON OUR WEBSITE.</ExplanationTxt>
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